subspecies

Lopaphus iolas iolas (Westwood, 1859)

Variable-Winged Lopaphus

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male, lateral view of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male: head & thorax, dorsal view of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male: head & thorax, lateral view of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male: end of abdomen, dorsal view of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559545; daab65d1-a490-4925-9187-c093a27b59df: copyright OUMNH. male: end of abdomen, lateral view of Necroscia iolas (holotype). (CollectionObject).

CollectionObject 1559552; NMW 899, 694f3d04-e0cc-4aaa-98fc-23e00bf9ab76: copyright NMW, Vienna. male of synonym Parasipyloidea subtilis (holotype). (CollectionObject).

Nomenclature (36)

Necroscia iolas Westwood, 1859: 145, pl. 19:2. (valid)
Holotype; male; daab65d1-a490-4925-9187-c093a27b59df; deposited at: Oxford University Museum of Natural History (OUM);
Candaules sparnius Stål, 1875: 87.
Syntypes (n= 2); female; 92380853-55ae-4bc4-b954-5039f431980f; deposited at: Naturhistorisches Museum, Wien (NMW);
... Show all ... (32)

Candaules sparnius Stål, 1875 (subjective synonym of Necroscia iolas Westwood, 1859) in Wood-Mason, 1877: 347.
Lopaphus iolas (Westwood, 1859) in Wood-Mason, 1877: 348.
Candaules iolas (Westwood, 1859) in Kirby, 1904: 367.
Candaules jolas [sic] (Westwood, 1859) in Redtenbacher, 1908: 539.
Parasipyloidea subtilis Redtenbacher, 1908: 480.
Holotype; female; NMW 899; deposited at: Naturhistorisches Museum, Wien (NMW); Perak
Candaules jolas [sic] (Westwood, 1859) (misspelling of Necroscia iolas Westwood, 1859) in Redtenbacher, 1908: 539.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, Brock & Seow-En, 1994: 8.
Parasipyloidea subtilis Redtenbacher, 1908 in Brock, 1995: 91.
Candaules sparnius Stål, 1875 (subjective synonym of Necroscia iolas Westwood, 1859) in Brock, 1995: 89.
Lopaphus iolas (Westwood, 1859) in Brock, 1995: 89.
Lopaphus iolas (Westwood, 1859) in Tay & Seow-Choen, 1996: 180.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 1997: 77, fig. 54.
Parasipyloidea subtilis Redtenbacher, 1908 in Brock, 1998: 61.
Lopaphus iolas (Westwood, 1859) in Rabenstein & Hausleithner, 1998: 1.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 1998: 89.
Parasipyloidea subtilis Redtenbacher, 1908 in Brock, 1999: 104.
Candaules jolas [sic] (Westwood, 1859) (synonym of Necroscia iolas Westwood, 1859) in Brock, 1999: 174.
Lopaphus iolas (Westwood, 1859) in Brock, 1999: 80, figs, 174.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 1999: 299, pl. 3e.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 1999: 75.
Lopaphus iolas (Westwood, 1859) in Seow-Choen & Goh, 1999: 264.
Parasipyloidea subtilis Redtenbacher, 1908 in Seow-Choen, 2000: 2, 27.
Candaules sparnius Stål, 1875 (subjective synonym of Necroscia iolas Westwood, 1859) in Otte & Brock, 2005: 186.
Parasipyloidea subtilis Redtenbacher, 1908 (subjective synonym of Necroscia iolas Westwood, 1859) in Otte & Brock, 2005: 186.
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 2005: 49.
Lopaphus iolas (Westwood, 1859) in Otte & Brock, 2005: 186.
Candaules sparnius Stål, 1875 (subjective synonym of Necroscia iolas Westwood, 1859) in Seow-Choen, 2017: 40. Phylogeny or classification
Parasipyloidea subtilis Redtenbacher, 1908 (subjective synonym of Necroscia iolas Westwood, 1859) in Seow-Choen, 2017: 40. Phylogeny or classification
Lopaphus iolas (Westwood, 1859) in Seow-Choen, 2017: 40, figs. Image or description; Phylogeny or classification
Lopaphus iolas (Westwood, 1859) in Rabihah, Lailatul-Nadhirah & Azman, 2018: 4. Specimen or distribution
Candaules sparnius Stål, 1875 (subjective synonym of Necroscia iolas Westwood, 1859) in Seow-Choen, 2021: 379. Phylogeny or classification
Parasipyloidea subtilis Redtenbacher, 1908 (subjective synonym of Necroscia iolas Westwood, 1859) in Seow-Choen, 2021: 379. Phylogeny or classification

Lopaphus iolas iolas (Westwood, 1859) in Seow-Choen, 2021: 379, figs. Image or description; Phylogeny or classification
Lopaphus iolas iolas (Westwood, 1859) in Brock & Büscher, 2022: 535. Specimen or distribution

Nomenclature references (23)

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html
Brock, P.D. (1995) Catalogue of Stick and Leaf-Insects (Insecta: Phasmida) associated with Peninsular Malaysia and Singapore. Malayan Nature Journal, 49(2), 83–102. Available at http://www.researchgate.net/publication/348052167_Catalogue_of_Stick_and_Leaf-Insects_Insecta_Phasmida_associated_with_Peninsular_Malaysia_and_Singapore_Malaysian_Naturalist_49283-102
... Show all ... (19)

Brock, P.D. (1998) Catalogue of type-specimens of Stick- and Leaf-Insects in the Naturhistorisches Museum Wien (Insecta: Phasmida). Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien: Vol. 13(5). Naturhistorischen Museum Wien, 72 pp. Available at http://www.zobodat.at/pdf/kat-nhmw_13_0003-0072.pdf

Brock, P.D. (1999) Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Kuala Lumpur, 223 pp. Available at http://www.nhbs.com/stick-and-leaf-insects-of-peninsular-malaysia-and-singapore-book

Kirby, W.F. (1904) A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). The Trustees of the British Museum, London, Vol. 1, 501 pp. Available at http://www.biodiversitylibrary.org/item/29661

Otte, D. & Brock, P.D. (2005) In Phasmida Species File. Catalog of Stick and Leaf Insects of the world. The Insect Diversity Association at the Academy of Natural Sciences, Philadelphia. 414 pp. Available at http://www.goodreads.com/book/show/24725959-phasmida-species-file

Rabenstein, R. & Hausleithner, B. (1998) Fressen und gefressen werden: Freilandbeobachtungen zur Ernährung und zu Prädatoren von Phasmiden im Gombak-Tal (W-Malaysia) (Phasmida: Heteronemiidae, Phasmatidae, Pseudophasmatidae). Mitt. Internat. Entomol. Ver. Frankfurt, 23(1/2), 1–12.

Rabihah, I., Lailatul-Nadhirah, A. & Azman, S. (2018) The diversity of stick insects (phasmatodea) on Gunung Jerai, Kedah. 1940(020053), 1–6. Available at http://www.researchgate.net/publication/324236928_The_diversity_of_stick_insects_phasmatodea_on_Gunung_Jerai_Kedah

Redtenbacher, J. (1908) Phasmidae Anareolatae (Phibalosomini, Acrophyllini, Necrosciini). In Die Insektenfamilie der Phasmiden: Vol. III. Verlag Engelmann, Leipzig. 339–589, pls 16–27. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmiden.Vol.Iii.PhasmidaeAnareolatae

Seow-Choen, F. & Goh, Y.Y. (1999) New records of stick insects from Pulau Tioman, Peninsular Malaysia, including description of a new species of Abrosoma (Phasmida: Pseudophasmatidae: Aschiphasmatinae). Raffles Bulletin of Zoology, Suppl. Available at http://rmbr.nus.edu.sg/rbz/biblio/s6/s6rbz263-269.pdf

Seow-Choen, F. (1997) A guide to the stick and leaf insects of Singapore. Singapore Science Centre, Singapore, 160 pp. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (1998) List of stick-insects (Phasmida) in the collection of the Centre for Insect Systematics Universiti Kebangsaan Malaysia. Serangga, 3(1), 87–92. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (1999) Stick and leaf insect (Phasmida: Insecta) biodiversity in the Nature Reserves of Singapore. Gardens´ Bulletin, Singapore, 49(2), 297–312. [1997] Available at http://www.biodiversitylibrary.org/page/43613590#page/317/mode/1up

Seow-Choen, F. (1999) The Phasmida. In C. Briffett & H.H. Chew (Eds.). State of the natural environment in Singapore. Nature Society, Singapore. pp. 68–76. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (2000) An Illustrated Guide to the Stick and Leaf Insects of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 173 pp. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (2005) Phasmids of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 120 pp. Available at http://www.nhpborneo.com/book/a-pocket-guide-phasmids-of-peninsular-malaysia-and-singapore/

Seow-Choen, F. (2017) A taxonomic guide to the stick insects of Singapore. Natural History Publications (Borneo), Kota Kinabalu, 163 pp. Available at http://www.nhpborneo.com/book/taxonomic-guide-stick-insects-singapore/

Seow-Choen, F. (2021) A taxonomic guide to the stick insects of Peninsular Malaysia: Vol. I. Natural History Publications (Borneo), Kota Kinabalu, 944 pp. Available at http://www.nhpborneo.com/a-taxonomic-guide-to-the-stick-insects-of-peninsular-malaysia/

Seow-Choen, F., Brock, P.D. & Seow-En, I. (1994) An introduction to the Stick and Leaf-insects of Singapore. Malaysian Naturalist, 48(1), 7–11. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Stål, C. (1875) Recensio orthopterorum. Revue critique des Orthoptères décrits par Linné, DeGeer et Thunberg. 3. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 32, 1–105. Available at http://hdl.handle.net/2027/coo.31924018284798

Tay, E.P. & Seow-Choen, F. (1996) Relationship of Plant families and stick-insects in Peninsular Malaysia and Singapore. In I.M. Turner, C.H. Diong, S.S.L. Lim, & P.K.L. Ng (Eds.). Biodiversity and the Dynamics of Ecosystems. DIWPA Series. Vol. 1, pp. 177–186. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Westwood, J.O. (1859) Catalogue of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. British Museum, London, 196 pp, 48 pl. Available at http://www.archive.org/details/catalogueofortho00brituoft
Wood-Mason, J. (1877) Notes on Phasmidae. Journal of the Asiatic Society Bengal, 46, 342–352, pl. 2–3. Available at http://www.biodiversitylibrary.org/item/110105#page/354/mode/1up

Descendants and synonyms

Stats

	Names
Rank	Total	Valid	Invalid
subspecies	7	1	6