Species

Anchiale austrotessulata Brock & Hasenpusch, 2007 ✕

Tessellated Stick-insect

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. mating pair, Broken Head, New South Wales, 8 February 2017. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. female kentomorphic phase, defence display, Maddock Wtlands, Beerwah, SE Queensland, 30 November 2011. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. female kentromorphic phase, Maddock Wtlands, Beerwah, SE Queensland, 30 November 2011. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. male, Maddock Wtlands, Beerwah, SE Queensland, 30 November 2011. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. female defence display, Maddock Wtlands, Beerwah, SE Queensland, 30 November 2011. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. male, Springbrook, SE Queensland, 4 February 2017. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. mating pair, Springbrook, SE Queensland, 4 February 2017. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. habitat, Springbrook, SE Queensland, 4 February 2017. (Otu).

Anchiale briareus (Gray, 1834): copyright Paul D. Brock. male, Cania, Queensland, 23 November 2011. (Otu).

Anchiale briareus (Gray, 1834): opyright Paul D. Brock. female, Cania, Queensland, 23 November 2011. (Otu).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female: head and thorax, lateral view of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female: end of abdomen, lateral view of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558209; b56337c7-e420-4e75-8dde-e36603b7f30d: copyright NHRS, Stockholm. male (paralectotype, left) and female (lectotype, on right) of synonym Ctenomorpha nigrovaria. (CollectionObject).

CollectionObject 1558211; MVMA T-13881, e31884a1-4c3d-40e5-91a9-1d11ddb56e21: copyright MVMA, Melbourne. male of synomyn Ctenomorpha tessulata (lectotype). (CollectionObject).

CollectionObject 1558212; NHMUK(SF IMPORT DUPLICATE) 845022, 464d8bf8-b5de-4bc4-a20e-1f290bc5ab0b: copyright Natural History Museum, London. male of synonym Ctenomorpha tessulata (paralectotype). (CollectionObject).

Nomenclature (131)

Phasma (Diura) briareus Gray, 1834: 45.
Holotype; female; MVMA T-13886; deposited at: Museum of Victoria (MVMA); Australia
Ctenomorpha macleayi Gray, 1835: 41.
Syntype; male; 1dd68a46-ca3c-459d-b6ee-4e0fd9acf8f9; deposited at: Linnean Society of London (LINN); Australia. Syntype; female; a9970be6-b7f9-4f18-b419-9f715f0b7cb9; deposited at: Linnean Society of London (LINN); Australia
... Show all ... (127)

Ctenomorpha tessulata Gray, 1835: 44. (homonym)
Lectotype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia
Acrophylla briareus (Gray, 1834) in Gray, 1835: 40.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Westwood, 1859: 115, pl. 35:1-2.
Syntype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia. Syntype; male; NHMUK(SF IMPORT DUPLICATE) 845022; deposited at: Natural History Museum, London (NHMUK); Australia
Acrophylla macleaii [sic] (Gray, 1835) in Westwood, 1859: 115.
Ctenomorpha tessellata [sic] (Westwood, 1859) (unjustified emendation for Ctenomorpha tessulata Gray, 1835)
Acrophylla macleaii [sic] (Gray, 1835) (misspelling of Ctenomorpha macleayi Gray, 1835) in Westwood, 1859: 115.
Acrophylla briareus (Gray, 1834) in Westwood, 1859: 114.
Ctenomorpha nigrovaria Stål, 1875: 83.
Lectotype; female; b56337c7-e420-4e75-8dde-e36603b7f30d; deposited at: Swedish Museum of Natural History, Entomology Collections (NHRS); Australia: Queensland: Queensland; Cape York
Acrophylla tessalata [sic] (Gray, 1835) in Rainbow, 1897: 41.
Ctenomorpha nigrovaria Stål, 1875 in Rainbow, 1897: 44.
Ctenomorpha macleayi Gray, 1835 (synonym of Acrophylla macleaii [sic] (Gray, 1835)) in Rainbow, 1897: 41.
Acrophylla tessalata [sic] (Gray, 1835) (misspelling of Ctenomorpha tessulata Gray, 1835) in Rainbow, 1897: 41.
Acrophylla briareus (Gray, 1834) in Rainbow, 1897: 40.
Anchiale tesselata [sic] Gray, 1835 in Tepper, 1903
Ctenomorpha nigrovaria Stål, 1875 in Tepper, 1903: 281. Specimen or distribution
Anchiale tesselata [sic] Gray, 1835 (misspelling of Ctenomorpha tessulata Gray, 1835) in Tepper, 1903
Acrophylla tesselata (Gray, 1835) in Tepper, 1903: 281. Image or description
Acrophylla macleayi (Gray, 1835) in Tepper, 1903: 281. Specimen or distribution
Acrophylla briareus (Gray, 1834) in Tepper, 1903: 280. Specimen or distribution
Ctenomorpha macleayi Gray, 1835 in Kirby, 1904: 389.
Ctenomorpha nigrovaria Stål, 1875 in Kirby, 1904: 389.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Kirby, 1904: 389.
Ctenomorpha briareus (Gray, 1834) in Kirby, 1904: 389.
Acrophylla macleai [sic] (Gray, 1835) in Redtenbacher, 1908: 458.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Redtenbacher, 1908: 458.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Redtenbacher, 1908: 459. Phylogeny or classification
Acrophylla macleai [sic] (Gray, 1835) (misspelling of Ctenomorpha macleayi Gray, 1835) in Redtenbacher, 1908: 458.
Ctenomorpha briareus (Gray, 1834) in Redtenbacher, 1908: 459, pl. 22:4.
Ctenomorphodes briareus (Gray, 1834) in Karny, 1923: 240.
Ctenomorpha briaria [sic] (Gray, 1834) in Sjöstedt, 1933: 6.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Sjöstedt, 1933: 6. Phylogeny or classification
Ctenomorpha briaria [sic] (Gray, 1834) (misspelling of Phasma (Diura) briareus Gray, 1834) in Sjöstedt, 1933: 6.
Ctenomorphodes tessulata (Gray, 1835) in Key, 1957: 247, pl. 1-3.
Ctenomorphodes tessulata (Gray, 1835) in Hadlington & Hoschke, 1959: 146. Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Key, 1960: 235-240.
Ctenomorphodes briareus (Gray, 1834) in Key, 1960: 236.
Acrophylla tessellata (Westwood, 1859) in Korboot, 1961: 161. Life history
Ctenomorphodes tessulatus (Gray, 1835) in Campbell, 1961: 112. Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Hadlington & Shipp, 1961: 268.
Ctenomorphodes briareus (Gray, 1834) in China & Riley, 1962: 19. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Heather, 1965: 33. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Heather, 1965: 86.
Ctenomorphodes tessulatus (Gray, 1835) in Campbell & Hadlington, 1967: 1. Life history; Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Key, 1970: 351, fig. 22.3g; 357-8. Image or description; Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Craddock, 1972: 448. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Campbell, 1974: 3. Life zone and ecology data
Ctenomorphodes tessulata (Gray, 1835) in Sellick, 1975: 1. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Rice & Webb, 1975: 277. Structure
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1976: 99.
Ctenomorphodes tessulatus (Gray, 1835) in Galloway, 1976: 446. Specimen or distribution
Ctenomorpha tessulatus Gray, 1835 in Bedford, 1978: 130.
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1980: 121, fig. 71. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Goode, 1980: 39. Specimen or distribution
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1980: 122, fig. 72. Image or description; Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in James, 1981: 5.
Ctenomorphodes tessulatus (Gray, 1835) in Hadlington & Johnson, 1982: 28. Specimen or distribution
Ctenomorpha macleayi Gray, 1835 in Vickery, 1983: 5.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Vickery, 1983: 5.
Ctenomorphodes briareus (Gray, 1834) in Vickery, 1983: 5.
Ctenomorphodes tessulata (Gray, 1835) in Vickery, 1983: 5. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Brock, 1985: 5, 30, figs. Image or description
Ctenomorphodes tessulata (Gray, 1835) in Carlberg, 1986: 1. Life history
Ctenomorphodes tessulata (Gray, 1835) in Carlberg, 1986: 643. Behavior
Ctenomorphodes tessulatus (Gray, 1835) in Masetti, 1987: 150. Structure
Ctenomorphodes tessulatus (Gray, 1835) in Carlberg, 1987: 48. Structure
Ctenomorphodes briareus (Gray, 1834) in Kevan, 1990: 27. Specimen or distribution
Ctenomorphodes tessulatus (Gray, 1835) in Preston-Mafham, 1990: 60. Image or description
Ctenomorphodes tessulatus (Gray, 1835) in Preston-Mafham & Preston-Mafham, 1993: 58. Image or description
Ctenomorpha tessellata [sic] (Westwood, 1859) in Crosby-Clark, 1995: 6. Specimen or distribution
Ctenomorphodes briareus (Gray, 1834) in Schulten, 1995: 85, figs., pl. 3.
Ctenomorphodes tessulata (Gray, 1835) in Rentz, 1996: 250, 255. Image or description
Ctenomorpha macleayi Gray, 1835 in Balderson, Rentz & Roach, 1998: 365.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Balderson, Rentz & Roach, 1998: 366.
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1998: 225. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Balderson, Rentz & Roach, 1998: 366.
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1998: 225. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1998: 280. Phylogeny or classification
Ctenomorphodes tessulata (Gray, 1835) in Balderson, Rentz & Roach, 1998: 366. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Seiler, Bradler & Koch, 2000: 59, figs.
Ctenomorphodes briareus (Gray, 1834) in Baxter, 2002: 44, figs.
Ctenomorphodes tessulata (Gray, 1835) in Baxter, 2002: 7. Image or description
Ctenomorphodes briareus (Gray, 1834) in Whiting, Bradler & Maxwell, 2003: 265.
Ctenomorphodes briareus (Gray, 1834) in Brock, 2003: 62.
Ctenomorphodes tessulata (Gray, 1835) in Brock, 2003: 62. Image or description; Life history
Ctenomorpha macleayi Gray, 1835 in Otte & Brock, 2005: 112.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Otte & Brock, 2005: 113.
Ctenomorphodes briareus (Gray, 1834) in Otte & Brock, 2005: 113.
Ctenomorphodes tessulata (Gray, 1835) in Otte & Brock, 2005: 113. Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hasenpusch & Brock, 2006: 4.
Anchiale austrotessulata Brock & Hasenpusch, 2007: 72.
Syntype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia. Syntype; male; NHMUK(SF IMPORT DUPLICATE) 845022; deposited at: Natural History Museum, London (NHMUK); Australia. Syntype; male; fe7926d8-498f-40d1-a7a2-c3e1615c6d96; deposited at: Oxford University Museum of Natural History (OUM); Australia
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2007: 12.
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Anchiale briareus (Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Anchiale briareus (Gray, 1834) in Brock, 2008: 12.
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Hasenpusch, 2009: 16, 96, figs., 165.
Anchiale briareus (Gray, 1834) in Buckley, Attanayake & Bradler, 2009: 1058. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Brock & Hasenpusch, 2009: 2, 16, 20, 97, figs., 165. Image or description; Phylogeny or classification; Specimen or distribution; Life history; Behavior
Anchiale briareus (Gray, 1834) in Harman, 2011: 8. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Harman, 2012: 16. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2012: 27, figs.. Life history; Behavior
Anchiale briareus (Gray, 1834) in Harman, 2012: 13. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Baker, 2015: 12. Ecological data
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Velonà, Brock, Hasenpusch & Mantovani, 2015: 458. Phylogeny or classification
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Hasenpusch, 2015: 51, figs. Image or description; Life history; Behavior
Acrophylla tessalata [sic] (Gray, 1835) (synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale tesselata [sic] Gray, 1835 (synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015 Phylogeny or classification
Ctenomorpha tessulata Gray, 1835 (objective synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Ctenomorpha tessellata [sic] (Westwood, 1859) (unjustified emendation for Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015 Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Velonà, Brock, Hasenpusch & Mantovani, 2015: 458. Phylogeny or classification
Ctenomorpha tessulata Gray, 1835 (objective synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Brock, Marshall, Beccaloni & Harman, 2016: 199. Phylogeny or classification; Specimen or distribution
Anchiale briareus (Gray, 1834) in Ripper, 2016: 16, fig.. Image or description
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Rudolf & Brock, 2017: e.T78796229A78796267. Ecological data; Specimen or distribution
Anchiale briareus (Gray, 1834) in Büscher, Buckley, Grohmann, Gorb & Bradler, 2018: 1-11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Rudolf & Brock, 2018: e.T78796358A78796497.
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2019: 19, figs. Specimen or distribution; Behavior
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Williams, 2020: 127. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Büscher, 2022: pl. 6, 453, 558, fig. Image or description; Specimen or distribution
Anchiale briareus (Gray, 1834) in Brock & Büscher, 2022: pl. 2, 9.3 & 13.2, 454, 558, figs. Image or description; Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Coupland, 2022: 39, figs. Image or description; Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Coupland, 2022: 5, fig. Image or description; Behavior
Anchiale austrotessulata Brock & Hasenpusch, 2007 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7-8. Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7. Phylogeny or classification

Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7-9. Image or description; Key; Specimen or distribution; Phylogeny or classification; Behavior

Nomenclature references (76)

Baker, E.W. (2015) The worldwide status of phasmids (Insecta: Phasmida) as pests of agriculture and forestry, with a generalised theory of phasmid outbreaks. Agriculture & Food Security, 4(22), 1–19. Available at http://agricultureandfoodsecurity.biomedcentral.com/articles/10.1186/s40066-015-0040-6
Balderson, J., Rentz, D.C.F. & Roach, A.M.E. (1998) Phasmatodea. In W.W.K. Houston & A. Wells (Eds.). Zoological Catalog of Australia. Vol. 23. Archaeognatha, Zygentoma, Blattodea, Isoptera, Mantodea, Dermaptera, Phasmatodea, Embioptera, Zoraptera. CSIRO Publishing, Melbourne. pp. 347–376, 451–456.
... Show all ... (72)

Baxter, R.N. (2002) Rearing Stick and Leaf Insects. Chudleigh Publishing, Ilford, 103 pp. Available at http://www.pemberleybooks.com/product/rearing-stick-and-leaf-insects/5624/

Bedford, G.O. (1978) Biology and ecology of the Phasmatodea. Annual Review of Entomology, 23, 125–149. Available at http://www.annualreviews.org/doi/abs/10.1146/annurev.en.23.010178.001013

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. & Coupland, R. (2022) Studying the phasmids of Springbrook Mountain, Binna Burra and surrounds, southeast Queensland. Phasmid Studies, 21, 15–74. Available at http://www.researchgate.net/publication/367341619_Studying_the_phasmids_of_Springbrook_Mountain_Binna_Burra_and_surrounds_southeast_Queensland

Brock, P.D. & Hasenpusch, J.W. (2007) Studies on the Australian stick insects (Phasmida), including a checklist of species and bibliography. Zootaxa, 1570, 1–84. Available at http://www.researchgate.net/publication/287837852_Studies_on_the_Australian_stick_insects_Phasmida_including_a_checklist_of_species_and_bibliography

Brock, P.D. & Hasenpusch, J.W. (2009) In The Complete Field Guide to Stick and Leaf Insects of Australia. CSIRO Publishing, Collingwood. 204 pp. Available at http://www.publish.csiro.au/pid/6012.htm

Brock, P.D. & Hasenpusch, J.W. (2015) Australian stick and leaf insects (Insecta, Phasmida): camouflage and natural history. In Camouflage cultures: beyond the art of disappearance. Sydney University Press, Sydney. pp. 47–64. Available at http://www.jstor.org/stable/j.ctt1bh4b60

Brock, P.D. (1985) In The Phasmid Rearer’s Handbook (Stick Insects and Leaf Insects). The Amateur Entomologists’ Society. The Amateur Entomologist Vol. 20, Feltham. 41 pp.

Brock, P.D. (2003) Rearing and studying stick and leaf insects. Revised Edition. The Amateur Entomologists' Society, London, Vol. 22, 89 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Brock, P.D. (2007) Exciting new Australian stick insects. Butterfly & Other Invertebrates Club Inc. Newsletter, 47, 11–12. Available at http://www.researchgate.net/profile/Paul-Brock-2

Brock, P.D. (2008) Exciting new Australian stick insects. The Phasmid Study Group Newsletter, 112, 10–12, front cover. [2007]

Brock, P.D. (2012) Kentromorphic phases in phasmids. The Phasmid Study Group Newsletter, 127, 27–28.

Brock, P.D. (2019) Kentromorphic phases in phasmids. The Phasmid Study Group Newsletter, 143, 19–20. Available at http://www.researchgate.net/publication/350837208_Kentromorphic_Phases_In_Phasmids

Brock, P.D., Marshall, J.A., Beccaloni, G.W. & Harman, A.J.E. (2016) The types of Phasmida in the Natural History Museum, London, UK. Zootaxa, 4179(2), 151–208. Available at http://www.researchgate.net/publication/309585799_The_types_of_Phasmida_in_the_Natural_History_Museum_London_UK

Buckley, T.R., Attanayake, D. & Bradler, S. (2009) Extreme convergence in stick insect evolution: phylogenetic placement of the Lord Howe Island tree lobster. Proceedings of the Royal Society B, 276, 1055–1062. [2008] Available at http://www.researchgate.net/publication/23763933_Extreme_convergence_in_stick_insect_evolution_Phylogenetic_placement_of_the_Lord_Howe_Island_tree_lobster

Büscher, T.H., Buckley, T.R., Grohmann, C., Gorb, S. & Bradler, S. (2018) The Evolution of Tarsal Adhesive Microstructures in Stick and Leaf Insects (Phasmatodea). Frontiers in Ecology and Evolution, 6, 69. Available at http://www.frontiersin.org/articles/10.3389/fevo.2018.00069/full

Campbell, K.G. & Hadlington, P. (1967) The biology of the three species of phasmatids which occur in plague numbers in forests of southeastern Australia. Forestry Commission of New South Wales Research Note, 20, 1–38. Available at http://www.dpi.nsw.gov.au/__data/assets/pdf_file/0005/390092/The-Biology-of-the-Three-Species-of-Phasmatids-Phasmatodea-which-occur-in-Plague-Numbers-in-Forests-of-Southeastern-Australia.pdf

Campbell, K.G. (1961) The effects of forest fires on three species of stick insects (Phasmatidae Phasmatodea) occurring in plagues in forest areas of south-eastern Australia. Proceedings of the Linnean Society of New South Wales, 86, 112–121. Available at http://www.biodiversitylibrary.org/page/34949471#page/118/mode/1up

Campbell, K.G. (1974) Factors limiting the distribution and abundance of the three species of phasmatids (Phasmatodea: Phasmatidae) which occur in plague numbers in forests of south-eastern Australia. Journal of the Entomological Society of Australia, 8, 3–6.

Carlberg, U. (1986) Evolutionary and ecological aspects of reproduction adaptations in Phasmida (Insecta). Biologisches Zentralblatt, 105, 641–651.

Carlberg, U. (1986) Phasmida - A biological review. Zoologischer Anzeiger, 216(1/2), 1–18.

Carlberg, U. (1987) Evolutionary and ecological aspects on ovarian diversity in Phasmida (Insecta). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 114, 45–63.

China, W.E. & Riley, N.D. (1962) Opinion 641. Addition of certain generic and specific names in the family Phasmatidae (Insecta, Phasmatodea) to the official lists and indexes. The Bulletin of Zoological Nomenclature, 19(5), 274–279. Available at http://www.biodiversitylibrary.org/page/12221157#page/318/mode/1up

Coupland, R. (2022) Observations on phasmid populations in Southeast Queensland, Australia following drought and subsequent rain events in the summer of 2019/2020. Phasmid Studies, 21, 4–14. Available at http://www.researchgate.net/publication/367339492_Observations_on_phasmid_populations_in_Southeast_Queensland_Australia_following_drought_and_subsequent_rain_events_in_the_summer_of_20192020

Craddock, E. (1972) Chromosonal diversity in the Australian Phasmatodea. Australian Journal of Zoology, 20, 445–462. Available at http://www.publish.csiro.au/zo/zo9720445

Crosby-Clark, S. (1995) A survey of the phasmids held in the collection of the National Museum of Wales, Cathays Park, Cardiff. Phasmid Studies, 4(1), 6–10. Available at http://www.biodiversitylibrary.org/item/207262#page/8/mode/1up

Galloway, I.D. (1976) Identifying Insects - Stick Insects. Queensland Agricultural Journal, 102, 445–446.

Goode, J. (1980) Stick and Leaf Insects. Order Phasmatodea (spectre or ghost insects). In Insects of Australia. Angus & Robertson Publishers, London. 39–42, pl. 1.

Gray, G.R. (1834) Descriptions of several species of Australian Phasmata. Transactions of the Entomological Society, London, 1, 45–46. Available at http://www.biodiversitylibrary.org/page/13496987/#page/63/mode/1up

Gray, G.R. (1835) Synopsis of the species of insects belonging to the family of Phasmidae. Longman, Rees, Orme, Brown, Green & Longman, London, 48 pp. Available at http://www.biodiversitylibrary.org/bibliography/8697

Hadlington, P. & Hoschke, F. (1959) Observations on the Ecology of the Phasmatid Ctenomorphodes tessulata Gray. Proceedings of the Linnean Society of New South Wales, 84, 146–159.

Hadlington, P. & Johnson, J.A. (1982) In An introduction to Australian insects. NSW University Press.

Hadlington, P. & Shipp, E. (1961) Diapause and parthenogenesis in the eggs of three species of Phasmatodea. Proceedings of the Linnean Society of New South Wales, 86(3), 268–279. Available at http://www.biodiversitylibrary.org/page/34949645/#page/292/mode/1up

Harman, A.J.E. (2011) The Development of the Phasmid Species List Part One: PSG No. 1-50. The Phasmid Study Group Newsletter, 125, 6–13. Available at http://phasmidstudygroup.org/files/PSG_Newsletters/Newsletter_125.pdf

Harman, A.J.E. (2012) The Development of the Phasmid Species List Part Four: PSG No.151 – PSG No.200. The Phasmid Study Group Newsletter, 129, 16–18.

Harman, A.J.E. (2012) The Development of the Phasmid Species List Part Two: PSG No. 51 - PSG No. 100. The Phasmid Study Group Newsletter, 127, 12–15.

Hasenpusch, J.W. & Brock, P.D. (2006) Studies on the Australian stick insect genus Ctenomorpha Gray (Phasmida: Phasmatidae), with the description of a new large species. Zootaxa, 1282, 1–15. Available at http://www.mapress.com/zootaxa/2006f/z01282p015f.pdf

Heather, N.W. (1965) Occurrence of Cleptidae (Hymenoptera) parasites in eggs of Ctenomorphodes tessulatus (Gray) (Phasmida, Phasmidae) in Queensland. Journal of the Entomological Society of Queensland, 4, 86. Available at http://onlinelibrary.wiley.com/doi/10.1111/j.1440-6055.1965.tb00662.x

Heather, N.W. (1965) Studies on female genitalia of Queensland Phasmida. Journal of the Entomological Society of Queensland, 4, 33–38. Available at http://onlinelibrary.wiley.com/doi/10.1111/j.1440-6055.1965.tb00643.x

Hennemann, F.H., Conle, O.V. & Suzuki, S. (2015) A study of the members of the tribe Phasmatini Gray, 1835, that occur within the boundaries of Wallacea (Phasmatodea: Phasmatidae: Phasmatinae: “Lanceocercata”). Zootaxa, 4008(1), 1–74. Available at http://www.researchgate.net/publication/281558622_Hennemann_FH_Conle_OC_Suzuki_S_2015_A_study_of_the_members_of_the_tribe_Phasmatini_Gray_1835_that_occur_within_the_boundaries_of_Wallacea_Phasmatodea_Phasmatidae_Phasmatinae_Lanceocercata_Zootaxa_4008

James, T. (1981) PSG/7 Ctenomorphodes briareus? The Phasmid Study Group Newsletter, 7, 5–6.

Jones, B.R., Brock, P.D., Coupland, R., Emmott, A. & Lo, N. (2026) A Revision of Australian Anchiale Stål, 1875 (Phasmida: Phasmatidae), with the Description of Two New Species. Austral Entomology, 65:e70096, 1–21. Available at https://onlinelibrary.wiley.com/doi/10.1111/aen.70096

Karny, H.H. (1923) Zur Nomenklatur der Phasmoiden. Hortus Botanicus Bogoriensis, Java (Treubia), 3(2), 230–242. Available at http://www.biodiversitylibrary.org/item/110723#page/310/mode/1up

Kevan, D.K.M.E. (1990) An annotated list of adventive and captive alien orthopteroid insects in Canada. Note, Lyman Entomological Museum and Research Laboratory, 16, 1–51.

Key, K.H.L. (1957) Kentromorphic phases in three species of Phasmatodea. Australian Journal of Zoology, 5(3), 247–284. Available at http://www.publish.csiro.au/zo/zo9570247

Key, K.H.L. (1960) Proposed addition of certain generic and specific names in the Family Phasmatidae (Class Insecta, Order Phasmatodea) to the official lists and indexes. Z.N.(S.) 1167. The Bulletin of Zoological Nomenclature, 17(6-8), 235–240. Available at http://www.biodiversitylibrary.org/page/12220208#page/265/mode/1uphttp://www.biodiversitylibrary.org/page/12220208#page/265/mode/1up

Key, K.H.L. (1970) In The Insects of Australia, A textbook for students and research workers. 1st Edition. Melbourne University Press, Melbourne. 348–359 pp.

Kirby, W.F. (1904) A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). The Trustees of the British Museum, London, Vol. 1, 501 pp. Available at http://www.biodiversitylibrary.org/item/29661

Korboot, K. (1961) Observations on life histories of Acrophylla tessellata Gray and Extatosoma tiaratum Macleay (Phasmida). University of Queensland Papers Department of Entomology, 1(2), 161–169.

Masetti, M. (1987) A comparative analysis of vitellins in stick insects. In M. Mazzini & V. Scali (Eds.). Stick Insects: Phylogeny and reproduction. Proceedings of the 1st International Symposium on Stick Insects, Siena. University of Siena. pp. 149–156. [1986]

Otte, D. & Brock, P.D. (2005) In Phasmida Species File. Catalog of Stick and Leaf Insects of the world. The Insect Diversity Association at the Academy of Natural Sciences, Philadelphia. 414 pp. Available at http://www.goodreads.com/book/show/24725959-phasmida-species-file

Preston-Mafham, K. (1990) In Grasshoppers and mantids of the world. Blandford, London.

Preston-Mafham, R. & Preston-Mafham, K. (1993) In The encyclopedia of land invertebrate behaviour. MIT Press. 1–320 pp.

Rainbow, W.J. (1897) Description of two new Australian Phasmas, together with a synopsis of the Phasmidae in Australia. Records of the Australian Museum, 3, 34–44, pls. 9–10. Available at http://www.biodiversitylibrary.org/page/6951489/#page/58/mode/1up

Redtenbacher, J. (1908) Phasmidae Anareolatae (Phibalosomini, Acrophyllini, Necrosciini). In Die Insektenfamilie der Phasmiden: Vol. III. Verlag Engelmann, Leipzig. 339–589, pls 16–27. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmiden.Vol.Iii.PhasmidaeAnareolatae

Rentz, D.C.F. (1996) Grasshopper country. The abundant orthopteroid insects of Australia. Orthoptera; grasshoppers, katydids, crickets. Blattodea; cockroaches. Mantodea; mantids. Phasmatodea; stick insects. University of New South Wales Press, Sydney, 284 pp. Available at http://books.google.co.uk/books?id=G0Z2wY2B2ncC&printsec=frontcover&dq=Grasshopper+country

Rice, M.J. & Webb, P.D. (1975) Giant axons in the central nervous system of Ctenomorphodes tessulatus (Gray), an Australian stick insect. Journal of the Australian Entomological Society, 14, 277–282. Available at http://onlinelibrary.wiley.com/doi/epdf/10.1111/j.1440-6055.1975.tb02042.x

Ripper, B. (2016) Searching for Stick insects in Queensland, Australia. The Phasmid Study Group Newsletter, 137, 16–18.

Rudolf, E. & Brock, P.D. (2017) Anchiale austrotessulata. Available at http://www.iucnredlist.org/details/78796229/0

Rudolf, E. & Brock, P.D. (2018) Anchiale briareus. p. e.T78796358A78796497. Available at http://www.iucnredlist.org/details/78796358/0

Schulten, D. (1995) In Wandelnde Blätter, Stab- und Gespenstschrecken. Entomologische Mitteilungen aus dem Löbbecke-Museum and Aquazoo. Vol. 3, 1–132 pp. [in German] Available at http://www.zobodat.at/pdf/Entomologie-heute_BH_3_0001-0132.pdf

Seiler, C., Bradler, S. & Koch, R. (2000) In Phasmiden Pflege und Zucht von Gespenstschrecken, Stabschrecken und Wandelnden Blattern im Terrarium. bede-Verlag GmbH, Ruhmannsfelden. 144 pp. [in German]

Sellick, J.T.C. (1975) The synonymy of phasmids. AES Exotic Entomology Group, 1975(3), 1.

Sellick, J.T.C. (1976) The eggs of stick insects (Phasmida): a review with descriptions of the eggs of eleven species. Systematic Entomology, 1, 95–105. Available at http://resjournals.onlinelibrary.wiley.com/doi/10.1111/j.1365-3113.1976.tb00342.x

Sellick, J.T.C. (1980) A study of the eggs of the insect order Phasmida with particular reference to establishing the taxonomic value of egg structure in this group. UNPUBLISHED. PhD thesis. University of London. Available at http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.281183

Sellick, J.T.C. (1998) The micropylar plate of the eggs of Phasmida, with a survey of the range of plate form within the order. Systematic Entomology, 23, 203–228. Available at http://onlinelibrary.wiley.com/doi/abs/10.1046/j.1365-3113.1998.00056.x

Sjöstedt, Y. (1933) Orthopterentypen im Naturhistorischen Reichsmuseum zu Stockholm. 7. Phasmidae. Arkiv för Zoologi, 25A(16), 1–10.

Stål, C. (1875) Recensio orthopterorum. Revue critique des Orthoptères décrits par Linné, DeGeer et Thunberg. 3. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 32, 1–105. Available at http://hdl.handle.net/2027/coo.31924018284798

Tepper, J.G.O. (1903) List of described genera and species of the Australian and Polynesian Phasmidae (Spectre-insects). Transactions of the Royal Society of Southern Australia, 26, 278–287. Available at http://www.biodiversitylibrary.org/item/136804#page/308/mode/1up

Velonà, A., Brock, P.D., Hasenpusch, J.W. & Mantovani, B. (2015) Cryptic diversity in Australian stick insects (Insecta; Phasmida) uncovered by the DNA barcoding approach. Zootaxa, 3957(4), 455–466. Available at http://www.mapress.com/zootaxa/2015/f/z03957p466f.pdf

Vickery, V.R. (1983) Catalogue of Australian stick insects (Phasmida, Phasmatodea, Phasmatoptera, or Cheleutoptera). CSIRO Australian division of Entomology, Technical Paper No 20, 1–19. Available at http://publications.csiro.au/rpr/download?pid=procite:bc6d4d28-0bab-4f42-8f59-c5025bb110cd&dsid=DS1

Westwood, J.O. (1859) Catalogue of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. British Museum, London, 196 pp, 48 pl. Available at http://www.archive.org/details/catalogueofortho00brituoft

Whiting, M.F., Bradler, S. & Maxwell, T. (2003) Loss and recovery of wings in stick insects. Priroda, 421(6920), 264–267. Available at http://whitinglab.byu.edu/Portals/61/docs/Publications/Nature_421.pdf
Williams, G. (2020) In The Invertebrate World of Australia’s Subtropical Rainforests. CSIRO Publishing, Clayton South, Vic. Available at http://www.publish.csiro.au/book/7948/

Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
Species	0	0	1	1