Species

Anchiale briareus (Gray, 1834)

Tessellated Stick-insect

Anchiale briareus (Gray, 1834): copyright J. Sztraka. large female nymph, Queensland: Kuranda, 12 January 2007. (Otu).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female: head and thorax, lateral view of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558207; MVMA T-13886, f81ef150-e846-4c69-bed7-16fa39ff323b: copyright MVMA, Melbourne. female: end of abdomen, lateral view of Phasma (Diura) briareus (holotype). (CollectionObject).

CollectionObject 1558209; b56337c7-e420-4e75-8dde-e36603b7f30d: copyright NHRS, Stockholm. male and female of synonym Ctenomorpha nigrovaria (syntypes). (CollectionObject).

CollectionObject 1558211; MVMA T-13881, e31884a1-4c3d-40e5-91a9-1d11ddb56e21: copyright MVMA, Melbourne. male of Ctenomorpha tessulata (syntype). (CollectionObject).

CollectionObject 1558212; NHMUK(SF IMPORT DUPLICATE) 845022, 464d8bf8-b5de-4bc4-a20e-1f290bc5ab0b: copyright Natural History Museum, London. male of Ctenomorpha tessulata (syntype). (CollectionObject).

Nomenclature (131)

Phasma (Diura) briareus Gray, 1834: 45.
Holotype; female; MVMA T-13886; deposited at: Museum of Victoria (MVMA); Australia
Ctenomorpha macleayi Gray, 1835: 41.
Syntype; male; 1dd68a46-ca3c-459d-b6ee-4e0fd9acf8f9; deposited at: Linnean Society of London (LINN); Australia. Syntype; female; a9970be6-b7f9-4f18-b419-9f715f0b7cb9; deposited at: Linnean Society of London (LINN); Australia
... Show all ... (127)

Ctenomorpha tessulata Gray, 1835: 44. (homonym)
Syntype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia. Syntype; male; NHMUK(SF IMPORT DUPLICATE) 845022; deposited at: Natural History Museum, London (NHMUK); Australia. Syntype; male; fe7926d8-498f-40d1-a7a2-c3e1615c6d96; deposited at: Oxford University Museum of Natural History (OUM); Australia
Acrophylla briareus (Gray, 1834) in Gray, 1835: 40.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Westwood, 1859: 115, pl. 35:1-2.
Syntype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia. Syntype; male; NHMUK(SF IMPORT DUPLICATE) 845022; deposited at: Natural History Museum, London (NHMUK); Australia. Syntype; male; fe7926d8-498f-40d1-a7a2-c3e1615c6d96; deposited at: Oxford University Museum of Natural History (OUM); Australia
Acrophylla macleaii [sic] (Gray, 1835) in Westwood, 1859: 115.
Ctenomorpha tessellata [sic] (Westwood, 1859) (unjustified emendation for Ctenomorpha tessulata Gray, 1835)
Acrophylla macleaii [sic] (Gray, 1835) (misspelling of Ctenomorpha macleayi Gray, 1835) in Westwood, 1859: 115.
Acrophylla briareus (Gray, 1834) in Westwood, 1859: 114.
Ctenomorpha nigrovaria Stål, 1875: 83.
Syntype; female; b56337c7-e420-4e75-8dde-e36603b7f30d; deposited at: Swedish Museum of Natural History, Entomology Collections (NHRS); Australia: Queensland: Queensland; Cape York. Syntype; male; f1c7b1ce-1061-4573-8d04-e32fc296579d; deposited at: Swedish Museum of Natural History, Entomology Collections (NHRS); Australia: Queensland: Queensland; Cape York
Acrophylla tessalata [sic] (Gray, 1835) in Rainbow, 1897: 41.
Ctenomorpha nigrovaria Stål, 1875 in Rainbow, 1897: 44.
Ctenomorpha macleayi Gray, 1835 (synonym of Acrophylla macleaii [sic] (Gray, 1835)) in Rainbow, 1897: 41.
Acrophylla tessalata [sic] (Gray, 1835) (misspelling of Ctenomorpha tessulata Gray, 1835) in Rainbow, 1897: 41.
Acrophylla briareus (Gray, 1834) in Rainbow, 1897: 40.
Anchiale tesselata [sic] Gray, 1835 in Tepper, 1903
Ctenomorpha nigrovaria Stål, 1875 in Tepper, 1903: 281. Specimen or distribution
Anchiale tesselata [sic] Gray, 1835 (misspelling of Ctenomorpha tessulata Gray, 1835) in Tepper, 1903
Acrophylla tesselata (Gray, 1835) in Tepper, 1903: 281. Image or description
Acrophylla macleayi (Gray, 1835) in Tepper, 1903: 281. Specimen or distribution
Acrophylla briareus (Gray, 1834) in Tepper, 1903: 280. Specimen or distribution
Ctenomorpha macleayi Gray, 1835 in Kirby, 1904: 389.
Ctenomorpha nigrovaria Stål, 1875 in Kirby, 1904: 389.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Kirby, 1904: 389.
Ctenomorpha briareus (Gray, 1834) in Kirby, 1904: 389.
Acrophylla macleai [sic] (Gray, 1835) in Redtenbacher, 1908: 458.
Ctenomorpha tessellata [sic] (Westwood, 1859) in Redtenbacher, 1908: 458.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Redtenbacher, 1908: 459. Phylogeny or classification
Acrophylla macleai [sic] (Gray, 1835) (misspelling of Ctenomorpha macleayi Gray, 1835) in Redtenbacher, 1908: 458.
Ctenomorpha briareus (Gray, 1834) in Redtenbacher, 1908: 459, pl. 22:4.
Ctenomorphodes briareus (Gray, 1834) in Karny, 1923: 240.
Ctenomorpha briaria [sic] (Gray, 1834) in Sjöstedt, 1933: 6.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Sjöstedt, 1933: 6. Phylogeny or classification
Ctenomorpha briaria [sic] (Gray, 1834) (misspelling of Phasma (Diura) briareus Gray, 1834) in Sjöstedt, 1933: 6.
Ctenomorphodes tessulata (Gray, 1835) in Key, 1957: 247, pl. 1-3.
Ctenomorphodes tessulata (Gray, 1835) in Hadlington & Hoschke, 1959: 146. Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Key, 1960: 235-240.
Ctenomorphodes briareus (Gray, 1834) in Key, 1960: 236.
Acrophylla tessellata (Westwood, 1859) in Korboot, 1961: 161. Life history
Ctenomorphodes tessulatus (Gray, 1835) in Campbell, 1961: 112. Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Hadlington & Shipp, 1961: 268.
Ctenomorphodes briareus (Gray, 1834) in China & Riley, 1962: 19. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Heather, 1965: 33. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Heather, 1965: 86.
Ctenomorphodes tessulatus (Gray, 1835) in Campbell & Hadlington, 1967: 1. Life history; Life zone and ecology data
Ctenomorphodes tessulatus (Gray, 1835) in Key, 1970: 351, fig. 22.3g; 357-8. Image or description; Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Craddock, 1972: 448. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Campbell, 1974: 3. Life zone and ecology data
Ctenomorphodes tessulata (Gray, 1835) in Sellick, 1975: 1. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Rice & Webb, 1975: 277. Structure
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1976: 99.
Ctenomorphodes tessulatus (Gray, 1835) in Galloway, 1976: 446. Specimen or distribution
Ctenomorpha tessulatus Gray, 1835 in Bedford, 1978: 130.
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1980: 121, fig. 71. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Goode, 1980: 39. Specimen or distribution
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1980: 122, fig. 72. Image or description; Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in James, 1981: 5.
Ctenomorphodes tessulatus (Gray, 1835) in Hadlington & Johnson, 1982: 28. Specimen or distribution
Ctenomorpha macleayi Gray, 1835 in Vickery, 1983: 5.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Vickery, 1983: 5.
Ctenomorphodes briareus (Gray, 1834) in Vickery, 1983: 5.
Ctenomorphodes tessulata (Gray, 1835) in Vickery, 1983: 5. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Brock, 1985: 5, 30, figs. Image or description
Ctenomorphodes tessulata (Gray, 1835) in Carlberg, 1986: 1. Life history
Ctenomorphodes tessulata (Gray, 1835) in Carlberg, 1986: 643. Behavior
Ctenomorphodes tessulatus (Gray, 1835) in Masetti, 1987: 150. Structure
Ctenomorphodes tessulatus (Gray, 1835) in Carlberg, 1987: 48. Structure
Ctenomorphodes briareus (Gray, 1834) in Kevan, 1990: 27. Specimen or distribution
Ctenomorphodes tessulatus (Gray, 1835) in Preston-Mafham, 1990: 60. Image or description
Ctenomorphodes tessulatus (Gray, 1835) in Preston-Mafham & Preston-Mafham, 1993: 58. Image or description
Ctenomorpha tessellata [sic] (Westwood, 1859) in Crosby-Clark, 1995: 6. Specimen or distribution
Ctenomorphodes briareus (Gray, 1834) in Schulten, 1995: 85, figs., pl. 3.
Ctenomorphodes tessulata (Gray, 1835) in Rentz, 1996: 250, 255. Image or description
Ctenomorpha macleayi Gray, 1835 in Balderson, Rentz & Roach, 1998: 365.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Balderson, Rentz & Roach, 1998: 366.
Ctenomorphodes briareus (Gray, 1834) in Sellick, 1998: 225. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Balderson, Rentz & Roach, 1998: 366.
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1998: 225. Phylogeny or classification
Ctenomorphodes tessulatus (Gray, 1835) in Sellick, 1998: 280. Phylogeny or classification
Ctenomorphodes tessulata (Gray, 1835) in Balderson, Rentz & Roach, 1998: 366. Phylogeny or classification
Ctenomorphodes briareus (Gray, 1834) in Seiler, Bradler & Koch, 2000: 59, figs.
Ctenomorphodes briareus (Gray, 1834) in Baxter, 2002: 44, figs.
Ctenomorphodes tessulata (Gray, 1835) in Baxter, 2002: 7. Image or description
Ctenomorphodes briareus (Gray, 1834) in Whiting, Bradler & Maxwell, 2003: 265.
Ctenomorphodes briareus (Gray, 1834) in Brock, 2003: 62.
Ctenomorphodes tessulata (Gray, 1835) in Brock, 2003: 62. Image or description; Life history
Ctenomorpha macleayi Gray, 1835 in Otte & Brock, 2005: 112.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Otte & Brock, 2005: 113.
Ctenomorphodes briareus (Gray, 1834) in Otte & Brock, 2005: 113.
Ctenomorphodes tessulata (Gray, 1835) in Otte & Brock, 2005: 113. Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hasenpusch & Brock, 2006: 4.
Anchiale austrotessulata Brock & Hasenpusch, 2007: 72.
Syntype; male; MVMA T-13881; deposited at: Museum of Victoria (MVMA); Australia. Syntype; male; NHMUK(SF IMPORT DUPLICATE) 845022; deposited at: Natural History Museum, London (NHMUK); Australia. Syntype; male; fe7926d8-498f-40d1-a7a2-c3e1615c6d96; deposited at: Oxford University Museum of Natural History (OUM); Australia
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2007: 12.
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Anchiale briareus (Gray, 1834) in Brock & Hasenpusch, 2007: 72.
Anchiale briareus (Gray, 1834) in Brock, 2008: 12.
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Hasenpusch, 2009: 16, 96, figs., 165.
Anchiale briareus (Gray, 1834) in Buckley, Attanayake & Bradler, 2009: 1058. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Brock & Hasenpusch, 2009: 2, 16, 20, 97, figs., 165. Image or description; Phylogeny or classification; Specimen or distribution; Life history; Behavior
Anchiale briareus (Gray, 1834) in Harman, 2011: 8. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Harman, 2012: 16. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2012: 27, figs.. Life history; Behavior
Anchiale briareus (Gray, 1834) in Harman, 2012: 13. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Baker, 2015: 12. Ecological data
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Velonà, Brock, Hasenpusch & Mantovani, 2015: 458. Phylogeny or classification
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Hasenpusch, 2015: 51, figs. Image or description; Life history; Behavior
Acrophylla tessalata [sic] (Gray, 1835) (synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale tesselata [sic] Gray, 1835 (synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015 Phylogeny or classification
Ctenomorpha tessulata Gray, 1835 (objective synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Ctenomorpha tessellata [sic] (Westwood, 1859) (unjustified emendation for Anchiale austrotessulata Brock & Hasenpusch, 2007) in Hennemann, Conle & Suzuki, 2015 Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Hennemann, Conle & Suzuki, 2015: 11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Velonà, Brock, Hasenpusch & Mantovani, 2015: 458. Phylogeny or classification
Ctenomorpha tessulata Gray, 1835 (objective synonym of Anchiale austrotessulata Brock & Hasenpusch, 2007) in Brock, Marshall, Beccaloni & Harman, 2016: 199. Phylogeny or classification; Specimen or distribution
Anchiale briareus (Gray, 1834) in Ripper, 2016: 16, fig.. Image or description
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Rudolf & Brock, 2017: e.T78796229A78796267. Ecological data; Specimen or distribution
Anchiale briareus (Gray, 1834) in Büscher, Buckley, Grohmann, Gorb & Bradler, 2018: 1-11. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Rudolf & Brock, 2018: e.T78796358A78796497.
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock, 2019: 19, figs. Specimen or distribution; Behavior
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Williams, 2020: 127. Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Büscher, 2022: pl. 6, 453, 558, fig. Image or description; Specimen or distribution
Anchiale briareus (Gray, 1834) in Brock & Büscher, 2022: pl. 2, 9.3 & 13.2, 454, 558, figs. Image or description; Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Brock & Coupland, 2022: 39, figs. Image or description; Specimen or distribution
Anchiale austrotessulata Brock & Hasenpusch, 2007 in Coupland, 2022: 5, fig. Image or description; Behavior
Anchiale austrotessulata Brock & Hasenpusch, 2007 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7-8. Phylogeny or classification
Ctenomorpha macleayi Gray, 1835 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7. Phylogeny or classification

Ctenomorpha nigrovaria Stål, 1875 (subjective synonym of Phasma (Diura) briareus Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7. Phylogeny or classification
Anchiale briareus (Gray, 1834) in Jones, Brock, Coupland, Emmott & Lo, 2026: 7-9. Image or description; Key; Specimen or distribution; Phylogeny or classification; Behavior

Nomenclature references (76)

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Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
Species	1	0	20	21