Root
Animalia
Polyneoptera
Orthopterida
Phasmida
Euphasmatodea
Phasmatidae
Phasmatinae
Phasmatini
Dryococelus
Dryococelus australis (Montrouzier, 1855)

species

Dryococelus australis (Montrouzier, 1855)

Lord Howe Island Stick-insect

Melbourne Zoo culture stock 2006. Egg: dorsal view. Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. male, lateral view (non type, Lord Howe Island). Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. female (non type, Lord Howe Island). Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. male (non type, Lord Howe Island). Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. male nymph: head and thorax, lateral view of synonym Eubulides spuria (possible holotype). Depicts CollectionObject 1558528; NHMUK(SF IMPORT DUPLICATE) 844951, e0b7f367-cf6b-49f5-b95a-5ec7d3ee494e, a CollectionObject.

copyright Natural History Museum, London. male nymph of synonym Eubulides spuria (possible holotype). Depicts CollectionObject 1558528; NHMUK(SF IMPORT DUPLICATE) 844951, e0b7f367-cf6b-49f5-b95a-5ec7d3ee494e, a CollectionObject.

Melbourne Zoo culture stock 2006. Female, dorsal view [Ball's Pyramid stock]. Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

Culture stock Melbourne Zoo 2006. Female nymph, dorsal view. Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. female, lateral view (non type, Lord Howe Island). Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. eggs (non type, Lord Howe Island). Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. male nymph, data labels of synonym Eubulides spuria (possible holotype). Depicts CollectionObject 1558528; NHMUK(SF IMPORT DUPLICATE) 844951, e0b7f367-cf6b-49f5-b95a-5ec7d3ee494e, a CollectionObject.

Egg showing operculum, Melbourne Zoo culture stock. Egg: view of operculum. Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

Melbourne Zoo culture stock 2006. Male, lateral view [Ball's Pyramid stock]. Depicts Dryococelus australis (Montrouzier, 1855), an Otu.

copyright Natural History Museum, London. male nymph, lateral view of synonym Eubulides spuria (possible holotype). Depicts CollectionObject 1558528; NHMUK(SF IMPORT DUPLICATE) 844951, e0b7f367-cf6b-49f5-b95a-5ec7d3ee494e, a CollectionObject.

Nomenclature (64)

Karabidion australe Montrouzier, 1855: 81. Image or description; Phylogeny or classification
Syntype; male; 563537b2-37df-40c2-9db0-e037388b67af; Australia: 'House-Island' [Lord Howe Island]. Syntype; female; da2663ca-468c-4974-9231-d20be56045d8; Australia: 'House-Island' [Lord Howe Island]
Eurycantha australis (Montrouzier, 1855) in Westwood, 1859: 65, pl. 1:1-2. Image or description; Phylogeny or classification
... Show all ... (60)

Karabidion australe Montrouzier, 1855 in Etheridge, 1889: 31. Specimen or distribution
Eurycantha australis (Montrouzier, 1855) in Rainbow, 1897: 38. Phylogeny or classification
Eurycantha (Karabidion) australis (Montrouzier, 1855) in Sharp, 1898: 274, fig. 160. Phylogeny or classification
Karabidion australis Montrouzier, 1855 in Tepper, 1903: 284. Specimen or distribution
Eubulides spuria Kirby, 1904: 441. Image or description; Phylogeny or classification
Holotype; immature; NHMUK(SF IMPORT DUPLICATE) 844951; deposited at: Natural History Museum, London (NHMUK); Brazil: Nova Hollandia
Eubulides spuria Kirby, 1904: 395. Phylogeny or classification
Karabidion australe Montrouzier, 1855 in Kirby, 1904: 395. Phylogeny or classification
Carabidion [sic] australe Montrouzier, 1855 in Redtenbacher, 1908: 340. Phylogeny or classification
Karabidion australe Montrouzier, 1855 in Lea, 1916: 145, pl. 9:17. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Gurney, 1947: 388, figs. 10-12. Image or description; Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Paramonov, 1963: 361. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in McAlpine, 1967: 71. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Smithers, 1970: 252. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Key, 1970: 358. Image or description; Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Healy & Smithers, 1971: 24. Ecological data
Dryococelus australis (Montrouzier, 1855) in Bedford, 1978: 140. Specimen or distribution
Eurycantha australis (Montrouzier, 1855) in Sellick, 1980: 112. Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Vickery, 1983: 6. Phylogeny or classification
Eubulides spuria Kirby, 1904 in Vickery, 1983: 6. Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Carlberg, 1986: 9. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Carwardine, 1995: 235. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Rentz, 1996: 249, figs., 256. Image or description
Dryococelus australis (Montrouzier, 1855) in Balderson, Rentz & Roach, 1998: 367. Phylogeny or classification
Eubulides spuria Kirby, 1904 in Balderson, Rentz & Roach, 1998: 367. Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Brock, 1999: 26, fig 14. Image or description
Dryococelus australis (Montrouzier, 1855) in Humphrey, 2001: 6. Image or description
Dryococelus australis (Montrouzier, 1855) in Priddel, Carlile, Humphrey, Fellenberg & Hiscox, 2003: 1391. Image or description; Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Otte & Brock, 2005: 130. Phylogeny or classification
Eubulides spuria Kirby, 1904 (subjective synonym of Karabidion australe Montrouzier, 1855) in Otte & Brock, 2005: 131.
Dryococelus australis (Montrouzier, 1855) in Cox, 2005: 8, figs.. Image or description
Dryococelus australis (Montrouzier, 1855) in Honan, 2006: 1. Life history
Dryococelus australis (Montrouzier, 1855) in Hennemann & Conle, 2006: 40, figs.. Phylogeny or classification
Eubulides spuria Kirby, 1904 (subjective synonym of Karabidion australe Montrouzier, 1855) in Hennemann & Conle, 2006: 42.
Dryococelus australis (Montrouzier, 1855) in Pain, 2006: 42. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Robertson, 2006: 781. Specimen or distribution
Eubulides spuria Kirby, 1904 (subjective synonym of Karabidion australe Montrouzier, 1855) in Brock & Hasenpusch, 2007: 72.
Dryococelus australis (Montrouzier, 1855) in Honan, 2007: 13, figs. Life history
Dryococelus australis (Montrouzier, 1855) in Honan, 2007: 258. Life history
Dryococelus australis (Montrouzier, 1855) in Honan, 2008: 399. Life history
Dryococelus australis (Montrouzier, 1855) in Henderson, Henderson & Sinclair, 2008: 86, fig.. Life history
Dryococelus australis (Montrouzier, 1855) in Buckley, Attanayake & Bradler, 2009: 1056, figs 1a-b. Image or description; Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Brock & Hasenpusch, 2009: 19, 35, 74-75 figs.. Image or description; Phylogeny or classification; Specimen or distribution; Life history; Behavior
Dryococelus australis (Montrouzier, 1855) in Carlile, Priddel & Honan, 2009: S124. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Egerton & Lochman, 2009: 367. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Priddel & Carlile, 2010: 17. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Lonsdale, 2010: 5. Ecological data
Dryococelus australis (Montrouzier, 1855) in Smith, 2012: 26, figs.. Image or description; Ecological data; Life history
Dryococelus australis (Montrouzier, 1855) in Smith, 2012: 35, figs.. Image or description; Ecological data; Life history
Dryococelus australis (Montrouzier, 1855) in Wilkinson, 2014: 1. Image or description; Ecological data; Life history
Dryococelus australis (Montrouzier, 1855) in Bradler, 2015: 1, fig.. Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Cleave & Tulloch, 2015: 1. Image or description; Ecological data
Eubulides spuria Kirby, 1904 (subjective synonym of Karabidion australe Montrouzier, 1855) in Brock, Marshall, Beccaloni & Harman, 2016: 197. Phylogeny or classification; Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Mikheyev, Zwick, Magrath, Grau, Qiu, Su & Yeates, 2017: 1. Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Smith, 2017: 16. Image or description; Phylogeny or classification
Dryococelus australis (Montrouzier, 1855) in Rudolf & Brock, 2017: e.T6852A21426226. Ecological data; Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Bower, Carlile, Cleave, Haselden, Hiscox & O'Neill, 2018: 189. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Bayley, Cheng & Lynch, 2018: 719. Behavior
Dryococelus australis (Montrouzier, 1855) in Flemons, Wills, Gray, Pearce, Priebbenow, Priebbenow, Mattick, Bell, Köhler, Bower, Hutton, Bannigan & Stephens, 2018: 77, figs. Specimen or distribution
Dryococelus australis (Montrouzier, 1855) in Smith, 2019: 6, figs. Economic matters
Dryococelus australis (Montrouzier, 1855) in Tiffin, 2019: 22. Specimen or distribution

Dryococelus australis (Montrouzier, 1855) in O'Hanlon, Jones & Bulbert, 2020: 2, fig. Image or description
Dryococelus australis (Montrouzier, 1855) in Brock & Büscher, 2022: 469, 558, figs. Image or description; Specimen or distribution

Nomenclature references (59)

Balderson, J., Rentz, D.C.F. & Roach, A.M.E. (1998) Phasmatodea. In W.W.K. Houston & A. Wells (Eds.). Zoological Catalog of Australia. Vol. 23. Archaeognatha, Zygentoma, Blattodea, Isoptera, Mantodea, Dermaptera, Phasmatodea, Embioptera, Zoraptera. CSIRO Publishing, Melbourne. pp. 347–376, 451–456.
Bayley, C., Cheng, C. & Lynch, M. (2018) Pathological and Microbiological Study of mortality in a Captive Breeding Colony of the Endangered Lord Howe Island Stick Insect (Dryococelus australis). 55(5), 719–730. Available at http://www.researchgate.net/publication/324568013_Pathological_and_Microbiological_Study_of_Mortality_in_a_Captive_Breeding_Colony_of_the_Endangered_Lord_Howe_Island_Stick_Insect_Dryococelus_australis
... Show all ... (55)

Bedford, G.O. (1978) Biology and ecology of the Phasmatodea. Annual Review of Entomology, 23, 125–149. Available at http://www.annualreviews.org/doi/abs/10.1146/annurev.en.23.010178.001013

Bower, H., Carlile, N., Cleave, R., Haselden, C., Hiscox, D. & O'Neill, L. (2018) The path to recovery for the ‘extinct’ Lord Howe Island phasmid Dryococelus australis. In Recovering Australian Threatened Species. CSIRO Publishing. pp. 189–198. Available at http://www.publish.csiro.au/book/7705/

Bradler, S. (2015) Der Phasmatodea Tree of Life: Überraschendes und Ungeklärtes in der Stabschrecken-Evolution [The Phasmatodea Tree of Life: Surprising Facts and Open Questions in the Evolution of Stick and Leaf Insects]. Entomologie heute, 27, 1–23. Available at http://www.zobodat.at/pdf/Entomologie-heute_27_0001-0023.pdf

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. & Hasenpusch, J.W. (2007) Studies on the Australian stick insects (Phasmida), including a checklist of species and bibliography. Zootaxa, 1570, 1–84. Available at http://www.researchgate.net/publication/287837852_Studies_on_the_Australian_stick_insects_Phasmida_including_a_checklist_of_species_and_bibliography

Brock, P.D. & Hasenpusch, J.W. (2009) In The Complete Field Guide to Stick and Leaf Insects of Australia. CSIRO Publishing, Collingwood. 204 pp. Available at http://www.publish.csiro.au/pid/6012.htm

Brock, P.D. (1999) In The amazing world of stick and leaf-insects. The Amateur Entomologists' Society, Orpington. Vol. 26, 1–165; 40 pp color plates. Available at http://shop.amentsoc.org/the-amazing-world-of-stick-and-leaf-insects.html

Brock, P.D., Marshall, J.A., Beccaloni, G.W. & Harman, A.J.E. (2016) The types of Phasmida in the Natural History Museum, London, UK. Zootaxa, 4179(2), 151–208. Available at http://www.researchgate.net/publication/309585799_The_types_of_Phasmida_in_the_Natural_History_Museum_London_UK

Buckley, T.R., Attanayake, D. & Bradler, S. (2009) Extreme convergence in stick insect evolution: phylogenetic placement of the Lord Howe Island tree lobster. Proceedings of the Royal Society B, 276, 1055–1062. [2008] Available at http://www.researchgate.net/publication/23763933_Extreme_convergence_in_stick_insect_evolution_Phylogenetic_placement_of_the_Lord_Howe_Island_tree_lobster

Carlberg, U. (1986) Phasmida - A biological review. Zoologischer Anzeiger, 216(1/2), 1–18.

Carlile, N., Priddel, D. & Honan, P. (2009) The recovery programme for the Lord Howe Island phasmid (Dryococelus australis) following its rediscovery. Ecological Management & Restoration, 10(Supp. 1), S124–S128. Available at http://www.researchgate.net/publication/230017970_The_recovery_programme_for_the_Lord_Howe_Island_Phasmid_Dryococelus_australis_following_its_rediscovery

Carwardine, M. (1995) In The Guinness Book of Animal Records. Guinness Publishing.

Cleave, R. & Tulloch, C. (2015) Phasmid: Saving the Lord Howe Island Stick Insect. CSIRO Publishing, Clayton, 33 pp. Available at http://www.publish.csiro.au/book/7226/

Cox, B. (2005) A trip to Melbourne. The Phasmid Study Group Newsletter, 102, 8–9.

Egerton, L. & Lochman, J. (2009) In Wildlife of Australia. Jacana Books, Allen & Unwin, Crows Nest. Available at http://books.google.co.uk/books/about/Wildlife_of_Australia.html?id=6ZbSAwAAQBAJ&redir_esc=y

Etheridge, R. (1889) The general zoology of Lord Howe Island. Australian Museum Memoirs, 2, 1–42. Available at http://journals.australian.museum/etheridge-1889-aust-mus-mem-21-142/

Flemons, P., Wills, V., Gray, D., Pearce, K., Priebbenow, Z., Priebbenow, P., Mattick, B., Bell, K., Köhler, F., Bower, H., Hutton, I., Bannigan, T. & Stephens, R. (2018) The Australian Museum Lord Howe Island Expedition 2017—Balls Pyramid Survey for Phasmids. Technical Reports of the Australian Museum, 26, 77–94. Available at http://doi.org/10.3853/j.1835-4211.26.2018.1708

Gurney, A.B. (1947) Notes on some remarkable Australasian walkingsticks, including a synopsis of the genus Extatosoma (Orthoptera: Phasmatidae). Annals of the Entomological Society of America, 40, 373–396. Available at http://academic.oup.com/aesa/article-abstract/40/3/373/18697

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Henderson, A., Henderson, D. & Sinclair, J. (2008) In Bugs alive: a guide to keeping Australian invertebrates. Museum Victoria, Melbourne. Available at http://shop.minibeastwildlife.com.au/bugs-alive-a-guide-to-keeping-australian-invertebrates/

Hennemann, F.H. & Conle, O.V. (2006) Papuacocelus papuanus n. gen., n. sp. — a new Eurycanthinae from Papua New Guinea, with notes on the genus Dryococelus Gurney, 1947 and description of the egg (Phasmatodea: Phasmatidae: Eurycanthinae). Zootaxa, 1375, 31–49. Available at http://www.researchgate.net/publication/263468601_Papuacocelus_Papuanus_N_Gen_N_Sp_A_New_Eurycanthinae_From_Papua_New_Guinea_With_Notes_On_The_Genus_Dryococelus_Gurney_1947_And_Description_Of_The_Egg_Phasmatodea_Phasmatidae_Eurycanthinae

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Honan, P. (2007) The Lord Howe Island Stick Insect: An example of the benefits of Captive Management. The Victorian Naturalist, 124(4), 258–261. Available at http://www.biodiversitylibrary.org/page/51254171#page/70/mode/1up

Honan, P. (2008) Notes on the biology, captive management and conservation status of the Lord Howe Island Stick Insect (Dryococelus australis) (Phasmatodea). Journal of Insect Conservation, 12, 399–413. Available at http://www.springerlink.com/content/95300q55k66g0757/

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Kirby, W.F. (1904) Notes on Phasmidae in the collection of the British Museum (Natural History), South Kensington, with descriptions of new species. - No. II. Annals and Magazine of Natural History, (7)13, 429–449. Available at http://www.tandfonline.com/doi/abs/10.1080/00222930408562475

Lea, A.M. (1916) Notes on the Lord Howe Island Phasma and on an associated longicorn beetle. Transactions of the Royal Society of Southern Australia, 40, 145–147, pl. 9–17.

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Mikheyev, A.S., Zwick, A., Magrath, M.J.L., Grau, M.L., Qiu, L., Su, Y.N. & Yeates, D. (2017) Museum Genomics Confirms that the Lord Howe Island Stick Insect Survived Extinction. Current Biology, 27, 1–5. Available at http://www.cell.com/current-biology/pdf/S0960-9822(17)31098-9.pdf

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O'Hanlon, J.C., Jones, B.R. & Bulbert, M.W. (2020) The dynamic eggs of the Phasmatodea and their apparent convergence with plants. The Science of Nature, 107(34), 1–12. Available at http://doi.org/10.1007/s00114-020-01690-1

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Smith, M. (2012) David Attenborough Travels Down Under to Meet Stick Insect Back From Extinction. The Phasmid Study Group Newsletter, 129, 35–36, figs.

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Smith, M. (2017) Confirmation that the Lord Howe Island Phasmid Survived Extinction. The Phasmid Study Group Newsletter, 139(16-17, figs).

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Tiffin, H. (2019) Many Happy Returns: Eradication, Re-Wilding and the Case of Lord Howe Island. Animal Studies Journal, 8(1/19), 8–29. Available at http://ro.uow.edu.au/cgi/viewcontent.cgi?article=1424&context=asj

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Wilkinson, R. (2014) Return of the Phasmid: Australia's rarest insect fights back from the brink of extinction. Media Dynamics, China, 150 pp.

Descendants and synonyms

Stats

	Names
Rank	Total	Valid	Invalid
species	7	1	6