species

Phyllium longicorne Latreille, 1802 ✕

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum head & thorax, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum end of abdomen, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum end of abdomen, ventral view (holotype). (CollectionObject).

CollectionObject 1590978; c1810eaa-1c0d-4ac2-8c10-4bf9e1713317: copyright MNHN, Paris. male (holotype). (CollectionObject).

Nomenclature (109)

Phyllium longicorne Latreille, 1802: 272.
Holotype; male; c1810eaa-1c0d-4ac2-8c10-4bf9e1713317; deposited at: Museum National d'Histoire Naturelle, Paris (MNHN); Unknown
Phyllium bioculatum Gray, 1832 in Griffith & Pidgeon, 1832: 191, pl. 63:3.
Holotype; male; 64124329-7811-4df4-829c-f138766ad5c3; deposited at: Oxford University Museum of Natural History (OUM); Unknown
... Show all ... (105)

Phyllium bioculatum Gray, 1832 in Gray, 1835: 30.
Phyllium bioculatum Gray, 1832 in Audinet-Serville, 1838: 292.
Phyllium bioculatum Gray, 1832 in Burmeister, 1838: 590.
Phyllium longicornis Latreille, 1802 in Guérin-Méneville, 1838: 414. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Gray, 1843: 117.
Phyllium bioculatum Gray, 1832 in Westwood, 1859: 175.
Phyllium bioculatum Gray, 1832 in Stål, 1875: 105.
Phyllium bioculatum Gray, 1832 in Bolívar, 1895: 374.
Phyllium bioculatum Gray, 1832 in Rehn, 1904: 106.
Pulchriphyllium bioculatum (Gray, 1832) in Kirby, 1904: 421.
Phyllium bioculatum Gray, 1832 in Redtenbacher, 1906: 174.
Pulchriphyllium bioculatum (Gray, 1832) in Giglio-Tos, 1910: 14.
Phyllium bioculatum Gray, 1832 in Bolívar & Ferrière, 1912: 294. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Merle, 1917: 93.
Phyllium bioculatum Gray, 1832 in Chopard, 1920: 261, 263. Phylogeny or classification; Image or description
Phyllium bioculatum Gray, 1832 in Grimpe, 1921: 227. Life history; Behavior
Pulchriphyllium bioculatum (Gray, 1832) in Showalter, 1929: 28, fig..
Phyllium bioculatum Gray, 1832 in Rehn & Rehn, 1934: 411.
Pulchriphyllium bioculatum (Gray, 1832) in Orian, 1957: 517. Specimen or distribution
Pulchriphyllium bioculatum (Gray, 1832) in Vinson, 1968: 113.
Phyllium bioculatum Gray, 1832 in Klante, 1969: 2.
Phyllium bioculatum Gray, 1832 in Sellick, 1974: 27.
Phyllium bioculatum Gray, 1832 in Bedford, 1978: 126. Life history
Phyllium bioculatum Gray, 1832 in Robinson, 1980: 2.
Phyllium bioculatum Gray, 1832 in Sellick, 1980: 197. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Vanschuytbroeck & Cools, 1981: 25. Specimen or distribution
Pulchriphyllium bioculatum (Gray, 1832) in Passerin d' Entrèves, 1981: 55. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Hausleithner, 1983: 278.
Phyllium bioculatum Gray, 1832 in Brock, 1985: 28, fig. Image or description
Phyllium bioculatum Gray, 1832 in Carlberg, 1986: 6.
Phyllium bioculatum Gray, 1832 in Carlberg, 1986: 643. Life history
Phyllium bioculatum Gray, 1832 in Carlberg, 1987: 54. Life history
Phyllium bioculatum Gray, 1832 in Carlberg, 1987: 47. Structure
Phyllium bioculatum Gray, 1832 in Matyot, 1988: 11.
Phyllium bioculatum Gray, 1832 in Sellick, 1988: 280. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Ziegler, 1989: 30.
Phyllium bioculatum Gray, 1832 in Flückiger & Peltier, 1989: figs.
Phyllium bioculatum Gray, 1832 in Matyot, 1990: 32.
Phyllium bioculatum Gray, 1832 in Matyot, 1990: 4. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Preston-Mafham, 1990: 82. Behavior
Phyllium bioculatum Gray, 1832 in Matyot, 1992: 18.
Phyllium bioculatum Gray, 1832 in Brock, 1992: 13, 46, fig.
Phyllium bioculatum Gray, 1832 in Unno, 1993: 12-13, fig. Image or description
Phyllium bioculatum Gray, 1832 in Seow-Choen, Brock & Seow-En, 1994: 8.
Phyllium bioculatum Gray, 1832 in Schulten, 1995: 108, figs., pl. 7.
Phyllium bioculatum Gray, 1832 in Deschandol, 1995: 20.
Phyllium bioculatum Gray, 1832 in Ziegler, 1995: 32.
Phyllium bioculatum Gray, 1832 in Woolman & Dharmasiri, 1995: 33. Ecological data
Phyllium bioculatum Gray, 1832 in Hanibeltz, Nakamura, Imms & Abdullah, 1995: 60. Life history
Phyllium bioculatum Gray, 1832 in Schulten, 1995: 643, fig. 4.
Phyllium bioculatum Gray, 1832 in Brock, 1995: 98.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 1995: 25, fig. Image or description
Phyllium bioculatum Gray, 1832 in Tay & Seow-Choen, 1996: 183.
Phyllium bioculatum Gray, 1832 in Seow-Choen, Seow-En & Seow-An, 1996: 43.
Phyllium bioculatum Gray, 1832 in Seiler, 1996: 6.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1996: 72, figs..
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1997: 139, figs..
Phyllium bioculatum Gray, 1832 in Alderton, 1997: 17, fig. Image or description
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1998: 90.
Phyllium bioculatum Gray, 1832 in Brock, 1999: 23, 41, pl. 18b.
Phyllium bioculatum Gray, 1832 in Brock, 1999: 160, pl. 35-38, 185.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1999: 299.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1999: 74.
Phyllium bioculatum Gray, 1832 in Brock, 2000: 56, figs.
Phyllium bioculatum Gray, 1832 in Seiler, Bradler & Koch, 2000: 110, figs.
Phyllium bioculatum Gray, 1832 in Delfosse, 2000: 3.
Phyllium bioculatum Gray, 1832 in Zompro, 2000: 61. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Brock, 2000: 9. Image or description
Phyllium bioculatum Gray, 1832 in Bragg, 2001: 191, fig..
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Grösser, 2001: 53, figs.
Phyllium bioculatum Gray, 1832 in Baxter, 2002: 14, 91, figs.
Phyllium bioculatum Gray, 1832 in Alderton, 2002: 16, fig. Image or description
Phyllium bioculatum Gray, 1832 in Whiting, Bradler & Maxwell, 2003: 265.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 2003: 226, fig. 1, 231.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 2003: 42, fig. 24.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Zompro, 2004: 273. Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2005: 108.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Otte & Brock, 2005: 275.
Phyllium bioculatum Gray, 1832 in Sorpongpaisal & Thanasinchayakul, 2006: 22. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Bragg, 2007: 9. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Gerlach & Haas, 2008: 68. Image or description; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Grösser, 2008: 93, figs.. Image or description; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Hennemann, Conle, Gottardo & Bresseel, 2009: 80. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Narita, Pereira, Kjellberg & Kageyama, 2010: 74. Structure
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Mandal & Yadav, 2010: 53. Image or description
Phyllium bioculatum Gray, 1832 in Harman, 2011: 7. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Hutacharern, 2011: 639. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Harman, 2012: 12-13. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Grösser, 2013: 46-51, figs.. Image or description; Phylogeny or classification; Specimen or distribution; Life history
Phyllium bioculatum Gray, 1832 in Bradler, 2015: 5, fig. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2016: 428, figs. Image or description
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2017: 153, figs. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2017: 202. Phylogeny or classification; Specimen or distribution
Phyllium bioculatum Gray, 1832 in Bradler & Buckley, 2018: 283, fig. Image or description
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Cumming, Le Tirant & Teemsma, 2018: 16. Image or description; Phylogeny or classification; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Cumming & Le Tirant, 2018: 2.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Laskar & Sinha, 2018: 1107-1111. Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2018: 666, figs. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, Seow-En, Chua & Choo, 2019: 382. Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2020: 397. Specimen or distribution
Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Bank, Cumming, Li, Henze, Le Tirant & Bradler, 2021: Supp. Table 1. Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Singh, Chandra & Gupta, 2021: 228. Specimen or distribution
Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Seow-Choen, 2021: 899, figs. Phylogeny or classification
Phyllium longicorne Latreille, 1802 in Brock & Büscher, 2022: 562. Specimen or distribution
Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Brock & Büscher, 2022: pl. 3-4 & 12, 425, 563, fig. Image or description; Specimen or distribution

Phyllium longicorne Latreille, 1802 (nomen oblitum of Phyllium bioculatum Gray, 1832) in Cumming, Foley, Hennemann, Le Tirant, Daawia, Warikar, Yando, Suhartawan, Henze, Büscher & Bank, 2025: 317, 321-323. Phylogeny or classification
Pulchriphyllium bioculatum (Gray, 1832) in Cumming, Foley, Hennemann, Le Tirant, Daawia, Warikar, Yando, Suhartawan, Henze, Büscher & Bank, 2025: 317, 321-323. Phylogeny or classification

Nomenclature references (107)

Alderton, D. (1997) Your first stick insect. Kingdom Books, Havant, 31 pp.
Alderton, D. (2002) How to Care for your Stick Insect. Kingdom Books, Havant, 34 pp.
... Show all ... (103)

Audinet-Serville, J.G. (1838) Histoire naturelle des Insectes. Orthoptères. Librarie Encyclopédique de Roret, Paris, 1–776, pls 1–14. [1839; in French] Available at http://www.archive.org/details/histoirenaturel01guilgoog

Bank, S., Cumming, R.T., Li, Y., Henze, K., Le Tirant, S. & Bradler, S. (2021) A tree of leaves: Phylogeny and historical biogeography of the leaf insects (Phasmatodea: Phylliidae). Communications Biology, 4(932), 1–12. Available at http://www.nature.com/articles/s42003-021-02436-z

Baxter, R.N. (2002) Rearing Stick and Leaf Insects. Chudleigh Publishing, Ilford, 103 pp. Available at http://www.pemberleybooks.com/product/rearing-stick-and-leaf-insects/5624/

Bedford, G.O. (1978) Biology and ecology of the Phasmatodea. Annual Review of Entomology, 23, 125–149. Available at http://www.annualreviews.org/doi/abs/10.1146/annurev.en.23.010178.001013

Bolívar, I. & Ferrière, C. (1912) Orthoptera, Phasmidae of the Seychelles. The Transactions of the Linnean Society of London, 15(2), 293–300. Available at http://www.biodiversitylibrary.org/page/16419304#page/337/mode/1up

Bolívar, I. (1895) Mission scientifique de M. Ch. Alluaud aux íles Séchelles (Mars, Avril, May, 1892). Orthoptères. Appendix Orthoptères de la Réunion 6 mémoire. Annales de la Société Entomologique de France, 64, 372–375. Available at http://www.biodiversitylibrary.org/item/34283#page/382/mode/1up

Bradler, S. & Buckley, T.R. (2018) 11. Biodiversity of Phasmatodea. In Insect Biodiversity: Science and Society. Volume 2. John Wiley & Sons Ltd. pp. 281–313. Available at http://www.researchgate.net/publication/326051925_Biodiversity_of_Phasmatodea

Bradler, S. (2015) Der Phasmatodea Tree of Life: Überraschendes und Ungeklärtes in der Stabschrecken-Evolution [The Phasmatodea Tree of Life: Surprising Facts and Open Questions in the Evolution of Stick and Leaf Insects]. Entomologie Heute, 27, 1–23. Available at http://www.zobodat.at/pdf/Entomologie-heute_27_0001-0023.pdf

Bragg, P.E. (2001) Phasmids of Borneo. Natural History Publications, Kota Kinabalu, Sabah, 771 pp. Available at http://www.nhpborneo.com/book/phasmids-of-borneo/

Bragg, P.E. (2007) Biographies of phasmatologists - 2. George Robert Gray. Phasmid Studies, 15(1&2), 5–9. Available at http://www.biodiversitylibrary.org/item/207388#page/9/mode/1up

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. (1985) In The Phasmid Rearer’s Handbook (Stick Insects and Leaf Insects). The Amateur Entomologists’ Society. The Amateur Entomologist Vol. 20, Feltham. 41 pp.

Brock, P.D. (1992) New Edition. Rearing and studying stick and leaf insects. The Amateur Entomologist. The Amateur Entomologists' Society, London, Vol. 22, 73 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Brock, P.D. (1995) An introduction to the world of stick & leaf insects. 25-27. Insect World International, 1(1), 25–27.

Brock, P.D. (1995) Catalogue of Stick and Leaf-Insects (Insecta: Phasmida) associated with Peninsular Malaysia and Singapore. Malayan Nature Journal, 49(2), 83–102. Available at http://www.researchgate.net/publication/348052167_Catalogue_of_Stick_and_Leaf-Insects_Insecta_Phasmida_associated_with_Peninsular_Malaysia_and_Singapore_Malaysian_Naturalist_49283-102

Brock, P.D. (1999) Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Kuala Lumpur, 223 pp. Available at http://www.nhbs.com/stick-and-leaf-insects-of-peninsular-malaysia-and-singapore-book

Brock, P.D. (1999) In The amazing world of stick and leaf-insects. The Amateur Entomologists' Society, Orpington. Vol. 26, 1–165; 40 pp color plates. Available at http://shop.amentsoc.org/the-amazing-world-of-stick-and-leaf-insects.html

Brock, P.D. (2000) A complete guide to breeding Stick and Leaf Insects. Kingdom Books, Havant, 1–67 pp.

Brock, P.D. (2000) Phasmid Postcards. The Phasmid Study Group Newsletter, 83, 9 & front cover.

Brock, P.D. (2003) Rearing and studying stick and leaf insects. Revised Edition. The Amateur Entomologists' Society, London, Vol. 22, 89 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Brock, P.D. (2003) Phasmida (Stick and leaf insects). In Grzimek's Animal Life Encyclopedia: Insects. Gale Group. pp. 221–231. Available at http://lemondedesphasmes.free.fr/IMG/pdf/sp657794.pdf

Burmeister, H.C.C. (1838) In Handbuch der Entomologie. T.C.F. Enslin, Berlin. Vol. 2, 457–756 pp. Available at http://www.biodiversitylibrary.org/bibliography/8135

Carlberg, U. (1986) Evolutionary and ecological aspects of reproduction adaptations in Phasmida (Insecta). Biologisches Zentralblatt, 105, 641–651.

Carlberg, U. (1986) Phasmida - A biological review. Zoologischer Anzeiger, 216(1/2), 1–18.

Carlberg, U. (1987) Culturing stick- and leaf-insects (Phasmida) - A review. Z. Versuchstierkd, 29, 39–63.

Carlberg, U. (1987) Evolutionary and ecological aspects on ovarian diversity in Phasmida (Insecta). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 114, 45–63.

Chopard, L. (1920) PhD thesis. Thèses Unicv. Paris, Rennes. In Recherches sur la conformation det le développement des derniers segments abdominaux chez le Orthoptères. 1–352 pp. [in French] Available at http://www.biodiversitylibrary.org/bibliography/50305#/summary

Cumming, R.T. & Le Tirant, S. (2018) Phyllium shurei n. sp., a third species of leaf insect from Java, Indonesia (Phasmida, Phylliidae). Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire, 6(1), 1–5. Available at http://www.researchgate.net/publication/322317383_Phyllium_shurei_n_sp_a_third_species_of_leaf_insect_from_Java_Indonesia_Phasmida_Phylliidae

Cumming, R.T., Foley, E.M., Hennemann, F.H., Le Tirant, S., Daawia, Warikar, E.L., Yando, H., Suhartawan, B., Henze, K., Büscher, T.H. & Bank, S. (2025) A deeper look into the diversity of Phyllium leaf insects from Indonesia: seven new species and two unique egg morphologies (Phasmatodea, Phylliidae). ZooKeys, 1256, 317–370. https://doi.org/10.3897/ zookeys.1256.162609

Cumming, R.T., Le Tirant, S. & Teemsma, S.N. (2018) On the Phyllium of Peninsular Malaysia and Sumatra, Indonesia, with range expansions for currently known species, description of the previously unknown Phyllium (Pu.) abdulfatahi Seow-Choen female, and description of the new species Phyllium (Ph.) rubrum n. sp. from Peninsular Malaysia (Phasmida: Phylliidae). Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire, 6(2), 1–21. Available at http://www.researchgate.net/publication/322901166_On_the_Phyllium_of_Peninsular_Malaysia_and_Sumatra_Indonesia_with_range_expansions_for_currently_known_species_description_of_the_previously_unknown_Phyllium_Pu_abdulfatahi_Seow-Choen_female_and_descr

Delfosse, E. (2000) Taxonomie, répartition, élevage et réflexions diverses notamment à propos du nom scientique du petit phasme réunionnais: Rhaphiderus scabrosus (Percheron, 1829) (Insecta Phasmatodea Phasmatidae Tropidoderinae Mandropterini). Bulletin de Phyllie, 4, 6–10.

Deschandol, A. (1995) Le mimétisme chez les Phyllium. Le Monde des Phasmes, 31(September), 20–22. Available at http://phasmidstudygroup.org/files/Le_Monde_Des_Phasmes/Le_Monde_des_Phasmes_31.pdf

Flückiger, P. & Peltier, M. (1989) In Gespenstschrecken. Naturmusems Olten.

Gerlach, J. & Haas, F. (2008) In Orthopteroidea of the Seychelles islands. Backhuys, Leiden, The Netherlands. Available at http://www.researchgate.net/publication/264166355_Orthopteroidea_of_the_Seychelles_Islands

Giglio-Tos, E. (1910) Fasmidi esotici del R. Museo zoologico di Torino e del Museo civico di Storia naturale di Genova. Bollettino dei Musei di Zoologia ed Anatomia comparata della Royal Università di Torino, 25(625), 1–57. Available at http://www.biodiversitylibrary.org/item/50963#page/57/mode/1up

Gray, G.R. (1835) Synopsis of the species of insects belonging to the family of Phasmidae. Longman, Rees, Orme, Brown, Green & Longman, London, 48 pp. Available at http://www.biodiversitylibrary.org/bibliography/8697

Gray, G.R. (1843) Description of several species of the genus Phyllium. Zoologist, 1(1), 117–123. Available at http://www.biodiversitylibrary.org/item/123076#page/137/mode/1up

Griffith, E. & Pidgeon, E. (1832) In The animal kingdom arranged in conformita with its organisation by the Baron Cuvier. Whittaker, London. Vol. 15, 1–796, pls 1–140.

Grimpe, G. (1921) Beiträge zur Biologie von Phyllium bioculatum G.R. Gray (Phasmidae). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 41, 227–266, pl. 13. Available at http://www.zobodat.at/pdf/Zoologische-Jahrbuecher-Syst_44_0227-0266.pdf

Grösser, D. (2001) In Wandelnde Blätter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen. 1st Edition. Edition Chimaira, Frankfurt am Main. 1–119 pp. [in German]

Grösser, D. (2008) In Wandelnde Blätter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen. 2nd Edition. Edition Chimaira, Frankfurt am Main. 1–175 pp. [in German] Available at http://www.nhbs.com/wandelnde-blatter-insekta-phasmidae-phylliidae-ein-katalog-aller-bisher-beschriebenen-phylliinae-arten-und-deren-eier-neubeschreibungen-book

Grösser, D. (2013) In Grundwissen Wandelnde Blätter eine Einführung in die Biologie, Haltung & Zucht. Sungaya-Verlag, Berlin. 1–79 pp. Available at http://sungaya-verlag.de/index.php/de/grundwissen

Guérin-Méneville, F.E. (1838) In Dictionnaire pittoresque d'histoire naturelle et des phénomènes de la nature. Paris. Vol. 7. Available at http://www.biodiversitylibrary.org/item/111205#page/559/mode/1up

Hanibeltz, A., Nakamura, Y., Imms, A. & Abdullah, E. (1995) The survival of newly-hatched leaf insects. Phasmid Studies, 4(2), 60–63. Available at http://www.biodiversitylibrary.org/item/207262#page/62/mode/1up

Harman, A.J.E. (2011) The Development of the Phasmid Species List Part One: PSG No. 1-50. The Phasmid Study Group Newsletter, 125, 6–13. Available at http://phasmidstudygroup.org/files/PSG_Newsletters/Newsletter_125.pdf

Harman, A.J.E. (2012) The Development of the Phasmid Species List Part Two: PSG No. 51 - PSG No. 100. The Phasmid Study Group Newsletter, 127, 12–15.

Hausleithner, B. (1983) Kritische Bemerkungen zu Phyllium bioculatum Gray und Ph. pulchrifolium Serville, zwei Vertreter der "wandelnden Blätter". (Phasmatodea: Phylliidae). Entomologische Zeitschrift, 93(19), 278–280.

Hennemann, F.H., Conle, O.V., Gottardo, M. & Bresseel, J. (2009) On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmatodea: Phylliidae: Phylliinae: Phylliini). Zootaxa, 2322, 1–83. Available at http://www.researchgate.net/publication/224856861_On_certain_species_of_the_genus_Phyllium_Illiger_1798_with_proposals_for_an_intra-generic_systematization_and_the_descriptions_of_five_new_species_from_the_Philippines_and_Palawan_Phasmatodea_Phylliid

Hutacharern, C. (2011) In Checklist of Insects in Thailand Book 2. Biological Diversity Division Office of Natural Resources and Environmental Policy and Planning Ministry of Natural Resources and Environment, Bangkok. Available at http://chm-thai.onep.go.th/doc/Publication/ONEP22/ONEP22-2.pdf

Kirby, W.F. (1904) A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). The Trustees of the British Museum, London, Vol. 1, 501 pp. Available at http://www.biodiversitylibrary.org/item/29661

Klante, H. (1969) Weitere Stabsneuschrecken (Insecta, Phasmatoptera) aus dem Naturkundemuseum Görlitz. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 44(7), 1–12.

Laskar, D. & Sinha, D.K. (2018) First report of Phyllium (Pulchriphyllium) bioculatum Gray, 1832 (Phylliidae: Phasmatodea): An addition to the fauna of Nagaland, North-East India. Journal of Entomology and Zoology Studies, 6(6), 1107–1111.

Latreille, P.A. (1802) Orthoptera. In Histoire naturelle, générale et particulière des crustacés et des insectes. Dufart, Paris. Vol. 3, 81–164, 270–274 pp. Available at http://www.biodiversitylibrary.org/item/80064#page/274/mode/1up

Mandal, S.K. & Yadav, K. (2010) Some phasmida (stick and leaf insects) of India. RECORDS OF THE ZOOLOGICAL SURVEY OF INDIA OCCASIONAL PAPER, 318, 1–64. Available at http://faunaofindia.nic.in/PDFVolumes/occpapers/318/index.pdf

Matyot, P. (1988) Phyllium bioculatum in the Seychelles. The Phasmid Study Group Newsletter, 37, 11.

Matyot, P. (1990) Notes on the leaf insect Phyllium bioculatum in the Seychelles. The Phasmid Study Group Newsletter, 42, 4.

Matyot, P. (1990) The Leaf Insect - master of disguise. Seychelles Today, 7, 32.

Matyot, P. (1992) Phasmids on Praslin and La Digue Islands in the Seychelles. Phasmid Studies, 1(1), 17–20. Available at http://www.biodiversitylibrary.org/item/207247#page/19/mode/1up

Merle, R. (1917) Les Phyllies Études biologiques de M. l'abbé Foucher. La Nature, 2263, 93–96.

Narita, S., Pereira, R.A.S., Kjellberg, F. & Kageyama, D. (2010) Gynandromorphs and intersexes: potential to understand the mechanism of sex determination in arthropods. Terrestrial Arthropod Reviews, 3, 63–96. Available at http://www.researchgate.net/publication/232274151_Narita_S_Pereira_RAS_Kjellberg_F_Kageyama_D_Gynandromorphs_and_intersexes_potential_to_understand_the_mechanism_of_sex_determination_in_arthropods_Terrestrial_Arthropod_Reviews

Orian, A.J.E. (1957) Saltatoria, Phasmida and Dictyoptera of Mauritius. Annals and Magazine of Natural History, 10(115), 513–520. Available at http://www.tandfonline.com/doi/abs/10.1080/00222935708655992

Otte, D. & Brock, P.D. (2005) In Phasmida Species File. Catalog of Stick and Leaf Insects of the world. The Insect Diversity Association at the Academy of Natural Sciences, Philadelphia. 414 pp. Available at http://www.goodreads.com/book/show/24725959-phasmida-species-file

Passerin d' Entrèves, P. (1981) Cataloghi. IV - Collezione Ortotterologiche del Museo di Zoologia de´ll Università di Torino. Museo Regionale di Scienze Naturali, Torino, 1–127 pp. Available at http://www.nhbs.com/1/series/cataloghi-di-museo-regionale-di-scienze-naturali-torino?qtview=185906

Preston-Mafham, K. (1990) In Grasshoppers and mantids of the world. Blandford, London.

Redtenbacher, J. (1906) Phasmidae Areolatae. In Die Insektenfamilie der Phasmiden. Vol. 1, 1–180, pls 1–6. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmidenVol.1

Rehn, J.A.G. & Rehn, J.W.H. (1934) On certain species of the genus Phyllium (Orthoptera; Phasmidae). Proceedings of the Academy of Natural Sciences of Philadelphia, 85, 411–427, pls. 16, 17. [1933] Available at http://www.jstor.org/stable/4064182?seq=1

Rehn, J.A.G. (1904) Studies in the Orthopterous family Phasmidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 56, 38–107. Available at http://www.biodiversitylibrary.org/item/30096#page/48/mode/1up

Robinson, D. (1980) Phyllium bioculatum (leaf insect) rearing. The Phasmid Study Group Newsletter, 3, 2.

Schulten, D. (1995) In Wandelnde Blätter, Stab- und Gespenstschrecken. Entomologische Mitteilungen aus dem Löbbecke-Museum and Aquazoo. Vol. 3, 1–132 pp. [in German] Available at http://www.zobodat.at/pdf/Entomologie-heute_BH_3_0001-0132.pdf

Schulten, D. (1995) Wandelnde Blätter, Stab- und Gespenstschrecken. Deutsche Aquarien und Terrarien Zeitschrift, 10/95, 642–646.

Seiler, C. (1996) Observations on the keeping and breeding of the walking leaf (Phyllium bioculatum, P. giganteum). The Phasmid Study Group Newsletter, 69, 6–7.

Seiler, C., Bradler, S. & Koch, R. (2000) In Phasmiden Pflege und Zucht von Gespenstschrecken, Stabschrecken und Wandelnden Blattern im Terrarium. bede-Verlag GmbH, Ruhmannsfelden. 144 pp. [in German]

Sellick, J.T.C. (1974) In Stick and Leaf Insects. Barry Shurlock, Winchester. 1–65 pp.

Sellick, J.T.C. (1980) A study of the eggs of the insect order Phasmida with particular reference to establishing the taxonomic value of egg structure in this group. UNPUBLISHED. PhD thesis. University of London. Available at http://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.281183

Sellick, J.T.C. (1988) The capitula of phasmid eggs: an update with a review of the current state of phasmid ootaxonomy. Zoological Journal of the Linnean Society of London, 93, 273–282. Available at http://academic.oup.com/zoolinnean/article-abstract/93/3/273/2658287?redirectedFrom=fulltext

Seow-Choen, F. (1996) The Leaf Insects of Peninsular Malaysia. Nature Malaysiana, 21(3), 68–73.

Seow-Choen, F. (1997) A guide to the stick and leaf insects of Singapore. Singapore Science Centre, Singapore, 160 pp. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (1998) List of stick-insects (Phasmida) in the collection of the Centre for Insect Systematics Universiti Kebangsaan Malaysia. Serangga, 3(1), 87–92. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (1999) Stick and leaf insect (Phasmida: Insecta) biodiversity in the Nature Reserves of Singapore. Gardens´ Bulletin, Singapore, 49(2), 297–312. [1997] Available at http://www.biodiversitylibrary.org/page/43613590#page/317/mode/1up

Seow-Choen, F. (1999) The Phasmida. In C. Briffett & H.H. Chew (Eds.). State of the natural environment in Singapore. Nature Society, Singapore. pp. 68–76. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (2005) Phasmids of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 120 pp. Available at http://www.nhpborneo.com/book/a-pocket-guide-phasmids-of-peninsular-malaysia-and-singapore/

Seow-Choen, F. (2016) A taxonomic guide to the stick insects of Borneo. Natural History Publications (Borneo), Kota Kinabalu, 454 pp. Available at http://www.nhpborneo.com/book/taxonomic-guide-stick-insects-borneo/

Seow-Choen, F. (2017) A taxonomic guide to the stick insects of Borneo: Vol. II. Natural History Publications (Borneo), Kota Kinabalu, Kota Kinabalu, 261 pp. Available at http://www.nhpborneo.com/book/a-taxonomic-guide-to-the-stick-insects-of-borneo-vol-2/

Seow-Choen, F. (2017) A taxonomic guide to the stick insects of Singapore. Natural History Publications (Borneo), Kota Kinabalu, 163 pp. Available at http://www.nhpborneo.com/book/taxonomic-guide-stick-insects-singapore/

Seow-Choen, F. (2018) A taxonomic guide to the stick insects of Sumatra: Vol. I. Natural History Publications (Borneo), Kota Kinabalu, 703 pp. Available at http://www.nhpborneo.com/book/a-taxonomic-guide-to-the-stick-insects-of-sumatra-vol-1/

Seow-Choen, F. (2020) A taxonomic guide to the stick insects of Sumatra: Vol. II. Natural History Publications (Borneo), Kota Kinabalu, 402 pp. Available at http://www.nhpborneo.com/books/animals/

Seow-Choen, F. (2021) A taxonomic guide to the stick insects of Peninsular Malaysia: Vol. I. Natural History Publications (Borneo), Kota Kinabalu, 944 pp. Available at http://www.nhpborneo.com/a-taxonomic-guide-to-the-stick-insects-of-peninsular-malaysia/

Seow-Choen, F., Brock, P.D. & Seow-En, I. (1994) An introduction to the Stick and Leaf-insects of Singapore. Malaysian Naturalist, 48(1), 7–11. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F., Seow-En, I. & Seow-An, S. (1996) Colour in stick & leaf insects. Nature Malaysiana, 21(2), 40–47. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F., Seow-En, I., Chua, E.K. & Choo, M.E. (2019) A Survey of Stick-Insects in Bukit Timah Nature Reserve, Singapore. 71(Suppl. 1), 381–389. Available at http://www.researchgate.net/publication/333397259_A_Survey_of_Stick-Insects_in_Bukit_Timah_Nature_Reserve_Singapore

Showalter, W.J. (1929) Exploring the wonders of the insect world. The National Geographic Magazine, 56(1), 26–28, pl.

Singh, L.R.K., Chandra, K. & Gupta, D. (2021) Insecta: Phasmida. In K. Chandra, L. Kosygin, C. Raghunathan, & D. Gupta (Eds.). Faunal Diversity of Biogeographic Zones of India: North-East. Zool. Surv. India, Kolkata. pp. 227–230.

Sorpongpaisal, W. & Thanasinchayakul, S. (2006) Identification of some Stick and Leaf Insects, (Order Phasmida) in Thailand. Kamphaengsean Academic Journal, 4(3), 18–32. [in Thai]

Stål, C. (1875) Recensio orthopterorum. Revue critique des Orthoptères décrits par Linné, DeGeer et Thunberg. 3. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 32, 1–105. Available at http://hdl.handle.net/2027/coo.31924018284798

Tay, E.P. & Seow-Choen, F. (1996) Relationship of Plant families and stick-insects in Peninsular Malaysia and Singapore. In I.M. Turner, C.H. Diong, S.S.L. Lim, & P.K.L. Ng (Eds.). Biodiversity and the Dynamics of Ecosystems. DIWPA Series. Vol. 1, pp. 177–186. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Unno, K. (1993) In Camouflage and Mimicry of Insects. Heibonsha Ltd. Publishers, Tokyo. 1–88 pp.

Vanschuytbroeck, P. & Cools, J. (1981) Catalogue et liste du material typique des Phasmatodea conserve dans les collections Entomologiques de l´Institut Royal des sciences naturelles de Belgique. Orthopteroidea: Phasmatodea Jacobson & Bianchi, 1902 (=Cheleutoptera Crompton, 1915). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 53(23), 1–26. Available at http://biblio.naturalsciences.be/rbins-publications/bulletins-de-linstitut-royal-des-sciences-naturelles-de-belgique-entomologie/bulletin-of-the-royal-belgian-institute-of-natural-sciences-entomology/vanschuytbroeck_catalogue_1981

Vinson, J. (1968) Les Blattopéroides, Orthopéroides et Dermaptéroides, des Mascareignes. The Mauritius Institute Bulletin, 6(3), 103–113.

Westwood, J.O. (1859) Catalogue of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. British Museum, London, 196 pp, 48 pl. Available at http://www.archive.org/details/catalogueofortho00brituoft

Whiting, M.F., Bradler, S. & Maxwell, T. (2003) Loss and recovery of wings in stick insects. Priroda, 421(6920), 264–267. Available at http://whitinglab.byu.edu/Portals/61/docs/Publications/Nature_421.pdf

Woolman, C. & Dharmasiri, H. (1995) Food plants for Phyllium bioculatum Gray in Sri Lanka. Phasmid Studies, 4(2), 33–36. Available at http://www.biodiversitylibrary.org/item/207262#page/35/mode/1up

Ziegler, U. (1989) Ein gynandromorphes Exemplar von Phyllium bioculatum (Gray) (Phasmida). Entomologische Zeitschrift, 99(3), 30–32.

Ziegler, U. (1995) Ein gynandromorphes Exemplar von Phyllium bioculatum (Gray) (Insecta: Phasmida). Arthropoda, 3(3), 32–33.

Zompro, O. (2000) Die Phasmidensammlung des Übersee-Museums Bremen. TenDenZen Supplement, 1999, 61–70. [1999] Available at http://www.researchgate.net/publication/27281173_Die_Phasmidensammlung_des_Ubersee_-_Museums_Bremen
Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Goecke & Evers, Keltern-Weiler, Germany, 327 pp. Available at http://www.researchgate.net/publication/313233330_Revision_of_the_genera_of_the_Areolatae_including_the_status_of_Timema_and_Agathemera_Insecta_Phasmatodea

Descendants and synonyms

Stats

	Valid names
Rank	Extant	Fossil	Invalid	Total
species	0	0	1	1