genus

Ramulus Saussure, 1862

Type

Bacillus (Ramulus) humberti Saussure, 1862 type species by original monotypy of Ramulus Saussure, 1862

Nomenclature (85)

Ramulus Saussure, 1862: 471. (valid)
type species by original monotypy Bacillus (Ramulus) humberti Saussure, 1862
Ramulus Saussure, 1862 in Saussure, 1869: 291. Phylogeny or classification
... Show all ... (81)

Clitumnus Stål, 1875: 8, 66.
type species by subsequent designation Phasma (Bacteria) nematodes Haan, 1842 in Kirby, 1904
Clitumnus Stål, 1875: 8. Phylogeny or classification
Paraclitumnus Brunner von Wattenwyl, 1893: 88, 91.
type species by subsequent designation Paraclitumnus lineatus Brunner von Wattenwyl, 1893 in Kirby, 1904
Paraclitumnus Brunner von Wattenwyl, 1893 in Kirby, 1904: 330.
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Kirby, 1904: 327. Phylogeny or classification
Ramulus Saussure, 1862 in Kirby, 1904: 329.
Clitumnus Stål, 1875 in Brunner von Wattenwyl, 1907: 187.
Dubreuilia Brunner von Wattenwyl, 1907: 208.
type species Paraclitumnus lineatus Brunner von Wattenwyl, 1893
Paraclitumnus Brunner von Wattenwyl, 1893 in Karny, 1923: 235.
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Karny, 1923: 235. Phylogeny or classification
Ramulus Saussure, 1862 in Karny, 1923: 236.
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Wu, 1935: 50. Phylogeny or classification
Dagys Günther, 1935: 3.
type species by original designation Dagys balia Günther, 1935
Ramulus Saussure, 1862 in Chopard, 1943: 94. Phylogeny or classification
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Günther, 1957: 87. Phylogeny or classification
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Sellick, 1975: 1. Phylogeny or classification
Dagys Günther, 1935 in Bradley & Galil, 1977: 185.
Paraclitumnus Brunner von Wattenwyl, 1893 in Bradley & Galil, 1977: 193.
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Bradley & Galil, 1977: 193. Phylogeny or classification
Ramulus Saussure, 1862 in Bradley & Galil, 1977: 185.
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Sellick, 1979: 327. Phylogeny or classification
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Sellick, 1980: 155. Phylogeny or classification
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1981: 31. Behavior
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1981: 568. Behavior
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1983: 127. Behavior
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1986: 11.
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1986: 39. Behavior
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1986: 511. Behavior
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Sellick, 1987: 137. Phylogeny or classification
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1987: 47. Structure
Ramulus Saussure, 1862 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Carlberg, 1987: 77. Structure
Ramulus Saussure, 1862 in Brock, 1991: 36.
Ramulus Saussure, 1862 in Brock, 1992: 13, 51. Image or description
Ramulus Saussure, 1862 in Brock & Shlagman, 1994: 104.
Ramulus Saussure, 1862 in Bragg, 1995: 12. Phylogeny or classification
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Balderson, Rentz & Roach, 1998: 363. Phylogeny or classification
Dagys Günther, 1935 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Hennemann, 1998: 124. Phylogeny or classification
Paraclitumnus Brunner von Wattenwyl, 1893 in Bragg, 2001: 639. Phylogeny or classification
Dubreuilia Brunner von Wattenwyl, 1907 (objective synonym of Paraclitumnus Brunner von Wattenwyl, 1893) in Bragg, 2001: 624.
Clitumnus Stål, 1875 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Bragg, 2001: 623. Phylogeny or classification
Dagys Günther, 1935 (subjective synonym of Bacillus (Baculum) Saussure, 1861) in Bragg, 2001: 623. Phylogeny or classification
Paraclitumnus Brunner von Wattenwyl, 1893 (subjective synonym of Ramulus Saussure, 1862) in Bragg, 2001: 639.
Ramulus Saussure, 1862 in Bragg, 2001: 626.
Clitumnus Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Hennemann, 2002: 64.
Ramulus Saussure, 1862 in Hennemann, 2002: 68.
Clitumnus Stål, 1875 in Zompro, 2004: 308.
Dagys Günther, 1935 in Zompro, 2004: 308.
Dubreuilia Brunner von Wattenwyl, 1907 in Zompro, 2004: 309.
Paraclitumnus Brunner von Wattenwyl, 1893 in Zompro, 2004: 317.
Ramulus Saussure, 1862 in Zompro, 2004: 321.
Clitumnus Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Otte & Brock, 2005: 299.
Dagys Günther, 1935 (subjective synonym of Ramulus Saussure, 1862) in Otte & Brock, 2005: 299.
Dubreuilia Brunner von Wattenwyl, 1907 (subjective synonym of Ramulus Saussure, 1862) in Otte & Brock, 2005: 299.
Paraclitumnus Brunner von Wattenwyl, 1893 (subjective synonym of Ramulus Saussure, 1862) in Otte & Brock, 2005: 299.
Ramulus Saussure, 1862 in Otte & Brock, 2005: 299.
Ramulus Saussure, 1862 in Brock & Hasenpusch, 2007: 73.
Paraclitumnus Brunner von Wattenwyl, 1893 in Chen & He, 2008: 220 [English 421].
Paraclitumnus Brunner von Wattenwyl, 1893 in Chen & He, 2008: 220.
Paraclitumnus Brunner von Wattenwyl, 1893 (subjective synonym of Ramulus Saussure, 1862) in Hennemann & Conle, 2008: 71.
Ramulus Saussure, 1862 in Hennemann & Conle, 2008: 71.
Ramulus Saussure, 1862 in Hennemann, Conle & Zhang, 2008: 26.
Ramulus Saussure, 1862 in Chen & He, 2008: 214 [English 420].
Ramulus Saussure, 1862 in Brock & Hasenpusch, 2009: 171. Image or description; Phylogeny or classification
Ramulus Saussure, 1862 in Mandal & Yadav, 2010: 25. Image or description
Ramulus Saussure, 1862 in Harman, 2012: 13. Specimen or distribution
Ramulus Saussure, 1862 in Harman, 2012: 16. Specimen or distribution
Ramulus Saussure, 1862 in Harman, 2013: 14. Specimen or distribution
Ramulus Saussure, 1862 in Ho, 2013: 73. Image or description; Phylogeny or classification
Ramulus Saussure, 1862 in Ho & Zhang, 2013: 525. Phylogeny or classification
Ramulus Saussure, 1862 in Harman, 2015: 27. Specimen or distribution
Ramulus Saussure, 1862 in Ichikawa, 2016: 189. Phylogeny or classification
Ramulus Saussure, 1862 in Ho, 2016: 330. Phylogeny or classification
Ramulus Saussure, 1862 in Seow-Choen, 2016: 262. Phylogeny or classification
Ramulus Saussure, 1862 in Seow-Choen, 2017: 92. Phylogeny or classification
Ramulus Saussure, 1862 in Seow-Choen, 2018: 398. Phylogeny or classification; Key
Ramulus Saussure, 1862 in Hennemann, 2021: 69, 171. Phylogeny or classification
Ramulus Saussure, 1862 in Seow-Choen, 2021: 707. Phylogeny or classification
Ramulus Saussure, 1862 in Brock & Büscher, 2022: 550. Specimen or distribution
Ramulus Saussure, 1862 in Harman, 2022: 21. Specimen or distribution
Ramulus Saussure, 1862 in Hennemann & Le Tirant, 2023: 12. Phylogeny or classification
Ramulus Saussure, 1862 in Seow-Choen, 2023: 422. Phylogeny or classification; Key

Ramulus Saussure, 1862 in Büscher, Boisseau & Brock, 2024: 449-450.
Ramulus Saussure, 1862 in Wang, Jeng, Tsai & Wu, 2025: 6-7. Phylogeny or classification

Nomenclature references (61)

Balderson, J., Rentz, D.C.F. & Roach, A.M.E. (1998) Phasmatodea. In W.W.K. Houston & A. Wells (Eds.). Zoological Catalog of Australia. Vol. 23. Archaeognatha, Zygentoma, Blattodea, Isoptera, Mantodea, Dermaptera, Phasmatodea, Embioptera, Zoraptera. CSIRO Publishing, Melbourne. pp. 347–376, 451–456.
Bradley, J.C. & Galil, B.S. (1977) The taxonomic arrangement of the Phasmatodea with keys to the subfamilies and tribes. Proceedings of the Entomological Society, Washington, 79(2), 176–208. Available at http://www.biodiversitylibrary.org/page/16301624#page/194/mode/1up
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Bragg, P.E. (1995) Type species of phasmid genera with particular reference to the status of Baculum Saussure, 1861, Ramulus, Saussure, 1862, and Gratidia Stål, 1875. Phasmid Studies, 4(1), 11–14. Available at http://www.biodiversitylibrary.org/item/207262#page/13/mode/1up

Bragg, P.E. (2001) Phasmids of Borneo. Natural History Publications, Kota Kinabalu, Sabah, 771 pp. Available at http://www.nhpborneo.com/book/phasmids-of-borneo/

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. & Hasenpusch, J.W. (2007) Studies on the Australian stick insects (Phasmida), including a checklist of species and bibliography. Zootaxa, 1570, 1–84. Available at http://www.researchgate.net/publication/287837852_Studies_on_the_Australian_stick_insects_Phasmida_including_a_checklist_of_species_and_bibliography

Brock, P.D. & Hasenpusch, J.W. (2009) In The Complete Field Guide to Stick and Leaf Insects of Australia. CSIRO Publishing, Collingwood. 204 pp. Available at http://www.publish.csiro.au/pid/6012.htm

Brock, P.D. & Shlagman, A. (1994) The stick-insects (Phasmatodea) of Israel, including the description of a new species. Israel Journal of Entomology, 28, 101–117. Available at http://www.entomology.org.il/sites/default/files/pdfs/IJE-1994.Brock_.pdf

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Brunner von Wattenwyl, C. (1893) Révision du Système des Orthoptères et description des espèces rapportées par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova, (2)13(33), 1–230. Available at http://www.biodiversitylibrary.org/bibliography/5121

Brunner von Wattenwyl, C. (1907) Phasmidae Anareolatae (Clitumnini, Lonchodini, Bacunculini). In Die Insektenfamilie der Phasmiden: Vol. II. Verlag Engelmann, Leipzig. 181–338, pls 7–15. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmiden.Vol.Ii.PhasmidaeAnareolataeclitumnini

Büscher, T.H., Boisseau, R.P. & Brock, P.D. (2024) Junior homonomy warrants a replacement name for Ramulus lineatus (Liu & Cai, 1992) (Phasmatodea: Phasmatidae). Zootaxa, 5447(3), 449–450. https://doi.org/10.11646/zootaxa.5447.3.12

Carlberg, U. (1981) Development period of eggs of a Baculum species (Phasmida., Phasmatidae). Entomologist’s monthly Magazine, 117, 31–34.

Carlberg, U. (1981) Storage of sperms in Phasmida (Insecta). Zoologische Jahrbücher Abteilung für Anatomie und Ontogenie der Tiere, 106, 568–571.

Carlberg, U. (1983) Diversity in the genus Baculum Saussure (Insecta: Phasmida). Zoologische Jahrbücher zur Systematik, 110, 127–140.

Carlberg, U. (1986) External genitalia ontogeny in Baculum sp. 1 (Insecta: Phasmida). Zoologische Jahrbücher Abteilung für Anatomie und Ontogenie der Tiere, 114, 511–516.

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Carlberg, U. (1986) Thanatosis and autotomy as defence in Baculum sp. 1 (Insecta: Phasmida). Zoologischer Anzeiger, 217(1/2), 39–53.

Carlberg, U. (1987) Evolutionary and ecological aspects on ovarian diversity in Phasmida (Insecta). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 114, 45–63.

Carlberg, U. (1987) Ovary anatomy in Phasmida (Insecta). Zoologische Jahrbücher Abteilung für Anatomie und Ontogenie der Tiere, 115, 77–84.

Chen, S.C. & He, Y.H. (2008) Phasmatodea of China. China Forestry Publishing House, Beijing, 476 pp. [in Chinese] Available at http://www.nhbs.com/phasmatodea-of-china-chinese-book

Chopard, L. (1943) Sous-ordre Phasmodea. In Orthoptéroïdes de l’Afrique du Nord. Faune de l’Empire Français. Lib. Larous, Paris. 88–95 pp.

Günther, K. (1935) Die von Gerd Heinrich 1930-32 auf Celebes gesammelten Phasmoiden. Mitteilungen aus dem Zoologischen Museum in Berlin, 21(1), 1–29, pl. 1–2.

Günther, K. (1957) Chapter III Phasmatoptera. In B. Hanstrom, P. Brinck, & G. Rudebeck (Eds.). South African Animal life, results of the Lund University expedition in 1950-51. Almqvist & Wiksell, Stockholm.

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Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Goecke & Evers, Keltern-Weiler, Germany, 327 pp. Available at http://www.researchgate.net/publication/313233330_Revision_of_the_genera_of_the_Areolatae_including_the_status_of_Timema_and_Agathemera_Insecta_Phasmatodea

Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
genus	1	0	5	6
species	150	0	303	453