species

Cladomorphus phyllinus Gray, 1835

Brazilian Giant Stick-insect

Cladomorphus petropolisensis Costa, Mallet & Takiya, 2022: copyright Jane Costa. Egg. (Otu).

CollectionObject 1557853; b47b1683-8fd3-4e67-a9dd-8de000215460: copyright NHRS, Stockholm. female juv. of synonym Phibalosoma coronata (holotype, on left). (CollectionObject).

CollectionObject 1557854; NHMUK(SF IMPORT DUPLICATE) 844926, 4b6d9ebd-7239-463f-b7aa-0c734b937214: copyright Natural History Museum, London. female juv., lateral view of synonym Cladomorphus dilatipes (holotype). (CollectionObject).

CollectionObject 1557857; f4b659c6-beed-4de5-886c-bb66ff39733e: from Piza 1938. female of synonym Phibalosoma rochai (holotype). (CollectionObject).

CollectionObject 1559457; NMW 558, 93c666d5-14d5-48af-8514-811677fe6fcb: copyright NMW, Vienna. female juv. of synonym Prisomera histrio (holotype). (CollectionObject).

CollectionObject 1559457; NMW 558, 93c666d5-14d5-48af-8514-811677fe6fcb: copyright NMW, Vienna. female juv. of synonym Prisomera histrio: head and thorax, lateral view (holotype). (CollectionObject).

CollectionObject 1559457; NMW 558, 93c666d5-14d5-48af-8514-811677fe6fcb: copyright NMW, Vienna. female juv. of synonym Prisomera histrio: end of abdomen, lateral view (holotype). (CollectionObject).

CollectionObject 1597679; 0c981c47-2724-4cb7-9f80-a067884b2742: copyright CEIOC Rio de Janeiro. female of synonym Cladomorphus petropolisensis, dorsal view (holotype). (CollectionObject).

CollectionObject 1597679; 0c981c47-2724-4cb7-9f80-a067884b2742: copyright CEIOC Rio de Janeiro. female head, lateral view of synonym Cladomorphus petropolisensis (holotype). (CollectionObject).

CollectionObject 1597679; 0c981c47-2724-4cb7-9f80-a067884b2742: copyright CEIOC Rio de Janeiro. female mesonotum of synonym Cladomorphus petropolisensis (holotype). (CollectionObject).

Nomenclature (86)

Cladomorphus phyllinus Gray, 1835: 15.
Syntype; ff9af12a-8588-448d-b4cc-cb69336af282; Brazil
Cladomorphus dilatipes Gray, 1835: 15.
Holotype; immature; NHMUK(SF IMPORT DUPLICATE) 844926; deposited at: Natural History Museum, London (NHMUK); Brazil
... Show all ... (82)

Phibalosoma lepelletieri Gray, 1835: 42.
Syntypes (n= 2); male; 480865ef-a4a0-4566-ac81-517f8c2cfa49; deposited at: Oxford University Museum of Natural History (OUM); Brazil
Cladomorphus phyllinus Gray, 1835 in Audinet-Serville, 1838: 225.
Cladomorphus phyllinus Gray, 1835 in Guérin-Méneville, 1838: 376, pl. 484. Image or description
Phibalosoma lepeletieri [sic] (Gray, 1835) in Audinet-Serville, 1838: 249.
Phibalosoma lepeletieri [sic] (Gray, 1835) (misspelling of Phibalosoma lepelletieri Gray, 1835) in Audinet-Serville, 1838: 249.
Cladoxerus phyllinus (Gray, 1835) in Burmeister, 1838: 571.
Cladomorphus phyllinus Gray, 1835 in Burmeister, 1840: 37.
Cladoxerus phyllinus (Gray, 1835) in Charpentier, 1845: pl. 34.
Phibalosoma lepeletierii [sic] (Gray, 1835) in Westwood, 1859: 72. Phylogeny or classification
Phibalosoma lepelletieri Gray, 1835 in Westwood, 1859: 72.
Cladomorphus dilatipes Gray, 1835 (synonym of Phibalosoma lepeletierii [sic] (Gray, 1835)) in Westwood, 1859: 72.
Phibalosoma lepeletierii [sic] (Gray, 1835) (misspelling of Phibalosoma lepelletieri Gray, 1835) in Westwood, 1859: 72.
Phibalosoma lepelieri [sic] (Gray, 1835) in Kaup, 1871: 36.
Phibalosoma lepelletieri Gray, 1835 (synonym of Phibalosoma lepeletieri [sic] (Gray, 1835)) in Kaup, 1871: 20, pl. 1:11a.
Phibalosoma lepelieri [sic] (Gray, 1835) (misspelling of Phibalosoma lepelletieri Gray, 1835) in Kaup, 1871: 36.
Phibalosoma coronata Stål, 1875: 80.
Holotype; immature; b47b1683-8fd3-4e67-a9dd-8de000215460; deposited at: Swedish Museum of Natural History, Entomology Collections (NHRS); Brazil
Cladomorphus phyllinas [sic] Gray, 1835 in Stål, 1875: 81.
Cladomorphus phyllinas [sic] Gray, 1835 (misspelling of Cladomorphus phyllinus Gray, 1835) in Stål, 1875: 81.
Cladoxerus phyllinus (Gray, 1835) in Göldi, 1886: 727, fig. 3-4a-c.
Phibalosoma coronata Stål, 1875 in Kirby, 1904: 357.
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Kirby, 1904: 356.
Cladoxerus phyllinus (Gray, 1835) in Rehn, 1904: 61.
Phibalosoma dilatipes (Gray, 1835) in Kirby, 1904: 356.
Phibalosoma phyllinus (Gray, 1835) in Kirby, 1904: 356.
Prisomera histrio Brunner von Wattenwyl, 1907: 287.
Holotype; immature; NMW 558; deposited at: Naturhistorisches Museum, Wien (NMW); Buru Island
Cladomorphus dilatipes Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Redtenbacher, 1908: 426.
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Redtenbacher, 1908: 426.
Phibalosoma phyllinum (Gray, 1835) in Redtenbacher, 1908: 426.
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Giglio-Tos, 1910: 42.
Phibalosoma phyllinum (Gray, 1835) in Giglio-Tos, 1910: 42.
Phibalosoma phyllinum (Gray, 1835) in Costa Lima, 1932: 258.
Lonchodes histrio (Brunner von Wattenwyl, 1907) in Günther, 1932: 380.
Phibalosoma coronata Stål, 1875 in Sjöstedt, 1933: 6.
Phibalosoma phyllinum (Gray, 1835) in Piza, 1938: 41.
Phibalosoma phyllinum (Gray, 1835) in Costa Lima, 1939: 189, fig. Specimen or distribution
Phibalosoma phyllinum (Gray, 1835) in Klante, 1969: 4.
Phibalosoma phyllinum (Gray, 1835) in Sellick, 1975: 1. Phylogeny or classification
Phibalosoma phyllinum (Gray, 1835) in Sellick, 1976: 101.
Phibalosoma phyllinum (Gray, 1835) in Sellick, 1980: 93. Phylogeny or classification
Phibalosoma phillinus [sic] (Gray, 1835) in Passerin d' Entrèves, 1981: 58.
Cladomorphus phyllium [sic] Gray, 1835 in Vanschuytbroeck & Cools, 1981: 13.
Phibalosoma phillinus [sic] (Gray, 1835) (misspelling of Cladomorphus phyllinus Gray, 1835) in Passerin d' Entrèves, 1981: 58.
Cladomorphus phyllium [sic] Gray, 1835 (misspelling of Cladomorphus phyllinus Gray, 1835) in Vanschuytbroeck & Cools, 1981: 13.
Phibalosoma phyllinum (Gray, 1835) in James, 1982: 7, figs.
Phibalosoma phyllinum (Gray, 1835) in Brock, 1985: 33, fig. Image or description
Phibalosoma phyllinum (Gray, 1835) in Brock, 1992: 13, 53, figs.
Phibalosoma phyllinum (Gray, 1835) in Hogue, 1993: 168, fig.
Phibalosoma phyllinum (Gray, 1835) in Sellick, 1998: 221. Phylogeny or classification
Lonchodes histrio (Brunner von Wattenwyl, 1907) in Brock, 1998: 33.
Cladomorphus phyllinus Gray, 1835 in Zompro, 2000: 66. Specimen or distribution
Cladomorphus phyllinus Gray, 1835 in Brock, 2003: 56.
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Zompro, 2004: 92.
Cladomorphus phyllinus Gray, 1835 in Otte & Brock, 2005: 95.
Phibalosoma coronata Stål, 1875 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Zompro & Domenico, 2005: 258.
Cladomorphus dilatipes Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Otte & Brock, 2005: 95.
Phibalosoma coronata Stål, 1875 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Otte & Brock, 2005: 95.
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Otte & Brock, 2005: 95.
Lonchodes histrio (Brunner von Wattenwyl, 1907) in Otte & Brock, 2005: 181.
Phibalosoma phyllinum (Gray, 1835) in Sottoriva, Picolo, Railda Roel & Hotta Ramos, 2007: 135. Life history
Cladomorphus phyllinus Gray, 1835 in Kumagai & Fonseca, 2009: 41. Phylogeny or classification
Cladomorphus phyllinus Gray, 1835 in Vargas, 2009: 1. Life history; Economic matters
Cladomorphus phyllinus Gray, 1835 in Harman, 2011: 7. Life history
Cladomorphus phyllinus Gray, 1835 in Azevedo, Fialho, Vargas, Vilela, Zanuncio & Serrão, 2013: 1417.
Cladomorphus phyllinus Gray, 1835 in Domínguez & Vera, 2014: 70. Specimen or distribution
Cladomorphus phyllinus Gray, 1835 in Shelomi, Sitepu, Boundy-Mills & Kimsey, 2015: 2. Structure
Cladomorphus phyllinus Gray, 1835 in Hennemann, Conle & Perez-Gelabert, 2016: 15, figs. Image or description
Cladomorphus dilatipes Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Brock, Marshall, Beccaloni & Harman, 2016: 168. Phylogeny or classification; Specimen or distribution
Cladomorphus phyllinus Gray, 1835 in Alvarenga, Souza, Giustolin, Matrangolo & Silva, 2018: 65. Life history
Cladomorphus phyllinus Gray, 1835 in Costa, Torres, Provance, Brugnera & Grazia, 2019: 5. Behavior
Cladomorphus phyllinus Gray, 1835 in Madeira-Ott, Thyssen & Costa, 2020: 2. Phylogeny or classification
Cladomorphus phyllinus Gray, 1835 in Brock & Büscher, 2022: 267, 546, figs. Image or description; Specimen or distribution
Lonchodes histrio (Brunner von Wattenwyl, 1907) in Brock & Büscher, 2022: 526. Specimen or distribution
Cladomorphus phyllinus Gray, 1835 in Costa, Mallet & Takiya, 2022: 4, figs. Image or description
Cladomorphus phyllinus Gray, 1835 in Torres, Benítez & Costa, 2022: 1. Behavior
Cladomorphus phyllinus Gray, 1835 in Costa, Chiquetto-Machado & Gil-Santana, 2022: 531. Behavior
Cladomorphus petropolisensis Costa, Mallet & Takiya, 2022: 9, figs.
Holotype; female; 0c981c47-2724-4cb7-9f80-a067884b2742; deposited at: Coleção Entomológica do Instituto Oswaldo Cruz (FIOCRUZ), Rio de Janeiro, Brazil (CEIOC); Brazil: Petrópolis
Cladomorphus phyllinus Gray, 1835 in Costa, Torres, Paschoaletto, Pimenta, Benítez, Suazo, Reigada & Gil-Santana, 2023: 1. Life history
Cladomorphus phyllinus Gray, 1835 in Heleodoro & Rafael, 2023: 14. Phylogeny or classification
Cladomorphus phyllinus Gray, 1835 in Hennemann & Conle, 2024: 46. Phylogeny or classification; Key
Cladomorphus petropolisensis Costa, Mallet & Takiya, 2022 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Hennemann & Conle, 2024: 46. Phylogeny or classification
Prisomera histrio Brunner von Wattenwyl, 1907 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Hennemann & Conle, 2024: 46. Phylogeny or classification
Cladomorphus dilatipes Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Hennemann & Conle, 2024: 46. Phylogeny or classification

Phibalosoma coronata Stål, 1875 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Hennemann & Conle, 2024: 46. Phylogeny or classification
Phibalosoma lepelletieri Gray, 1835 (subjective synonym of Cladomorphus phyllinus Gray, 1835) in Hennemann & Conle, 2024: 46. Phylogeny or classification

Nomenclature references (57)

Alvarenga, C.D., Souza, H.R., Giustolin, T.A., Matrangolo, C.A.R. & Silva, J.F. (2018) Biologia de Cladomorphus phyllinus Gray (Phasmatodea: Phasmatidae) em folhas de goiabeira (Psidium guajava). Entomobrasilis, 11(2), 65–69.
Audinet-Serville, J.G. (1838) Histoire naturelle des Insectes. Orthoptères. Librarie Encyclopédique de Roret, Paris, 1–776, pls 1–14. [1839; in French] Available at http://www.archive.org/details/histoirenaturel01guilgoog
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Azevedo, D.O., Fialho, M.D.C.Q.F., Vargas, N.C., Vilela, E.F., Zanuncio, J.C. & Serrão, J.E. (2013) Morphology of the digestive tract of Cladomorphus phyllinus (Phasmatodea: Phasmidae). The Florida Entomologist, 96(4), 1417–1423. Available at http://bioone.org/journals/florida-entomologist/volume-96/issue-4/024.096.0421/Morphology-of-the-Digestive-Tract-of-Cladomorphus-phyllinus-Phasmatodea/10.1653/024.096.0421.full

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. (1985) In The Phasmid Rearer’s Handbook (Stick Insects and Leaf Insects). The Amateur Entomologists’ Society. The Amateur Entomologist Vol. 20, Feltham. 41 pp.

Brock, P.D. (1992) New Edition. Rearing and studying stick and leaf insects. The Amateur Entomologist. The Amateur Entomologists' Society, London, Vol. 22, 73 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Brock, P.D. (1998) Catalogue of type-specimens of Stick- and Leaf-Insects in the Naturhistorisches Museum Wien (Insecta: Phasmida). Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien: Vol. 13(5). Naturhistorischen Museum Wien, 72 pp. Available at http://www.zobodat.at/pdf/kat-nhmw_13_0003-0072.pdf

Brock, P.D. (2003) Rearing and studying stick and leaf insects. Revised Edition. The Amateur Entomologists' Society, London, Vol. 22, 89 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Brock, P.D., Marshall, J.A., Beccaloni, G.W. & Harman, A.J.E. (2016) The types of Phasmida in the Natural History Museum, London, UK. Zootaxa, 4179(2), 151–208. Available at http://www.researchgate.net/publication/309585799_The_types_of_Phasmida_in_the_Natural_History_Museum_London_UK

Brunner von Wattenwyl, C. (1907) Phasmidae Anareolatae (Clitumnini, Lonchodini, Bacunculini). In Die Insektenfamilie der Phasmiden: Vol. II. Verlag Engelmann, Leipzig. 181–338, pls 7–15. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmiden.Vol.Ii.PhasmidaeAnareolataeclitumnini

Burmeister, H.C.C. (1838) In Handbuch der Entomologie. T.C.F. Enslin, Berlin. Vol. 2, 457–756 pp. Available at http://www.biodiversitylibrary.org/bibliography/8135

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Costa Lima, A.M.da (1939) XI. Ordem Phasmida. In Insectos do Brasil. 1' Tomo. Esc. Nac. Agr., Rio de Janeiro. 189–204 pp. Available at http://www.ephemeroptera-galactica.com/pubs/pub_c/pubcostaa1938p1.pdf

Costa, J., Chiquetto-Machado, P.I. & Gil-Santana, H.R. (2022) Predation strategies of Harpactor angulosus (Lepeletier & Serville, 1825) (Hemiptera: Reduviidae) on Cladomorphus phyllinus Gray, 1835 (Phasmatodea: Phasmatidae) in captivity. Revista Brasileira de Entomologia, 48(3), 531–543. Available at http://www.researchgate.net/publication/362336376_Predation_strategies_of_Harpactor_angulosus_Lepeletier_Serville_1825_Hemiptera_Reduviidae_on_Cladomorphus_phyllinus_Gray_1835_Phasmatodea_Phasmatidae_in_captivity

Costa, J., Mallet, J.R.S. & Takiya, D.M. (2022) Cladomorphus petropolisensis, a New Species of Stick Insect from the Atlantic Forest, Rio de Janeiro, Brazil. Animals, 12(2871), 1–14. Available at http://www.mdpi.com/2076-2615/12/20/2871

Costa, J., Torres, L., Paschoaletto, L., Pimenta, A.L.A., Benítez, H.A., Suazo, M.J., Reigada, C. & Gil-Santana, H.R. (2023) Unraveling the Sexual Dimorphism of First Instar Nymphs of the Giant Stick Insect, Cladomorphus phyllinus Gray, 1835, from the Atlantic Forest, Brazil. Animals, 13(3474), 1–13. Available at https:// doi.org/10.3390/ani13223474

Costa, J., Torres, L., Provance, D.W., Brugnera, R. & Grazia, J. (2019) First report of predation by a stink bug (Supputius cincticeps Stål) on a walking-stick insect (Cladomorphus phyllinus Gray), with reflections on evolutionary mechanisms for camouflage. Acta Biologica Paranaense, Curitiba, 48(1-2), 5–15.

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Gray, G.R. (1835) Synopsis of the species of insects belonging to the family of Phasmidae. Longman, Rees, Orme, Brown, Green & Longman, London, 48 pp. Available at http://www.biodiversitylibrary.org/bibliography/8697

Guérin-Méneville, F.E. (1838) In Dictionnaire pittoresque d'histoire naturelle et des phénomènes de la nature. Paris. Vol. 7. Available at http://www.biodiversitylibrary.org/item/111205#page/559/mode/1up

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Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
species	1	0	16	17