species

Pulchriphyllium bioculatum (Gray, 1832)

Javanese Leaf Insect; Gray's Leaf Insect; Javanese Leaf-insect; Gray's Leaf-insect

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum head & thorax, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum end of abdomen, dorsal view (holotype). (CollectionObject).

CollectionObject 1561239; 64124329-7811-4df4-829c-f138766ad5c3: copyright UMO. male Phyllium bioculatum end of abdomen, ventral view (holotype). (CollectionObject).

Nomenclature (104)

Phyllium bioculatum Gray, 1832 in Griffith & Pidgeon, 1832: 191, pl. 63:3.
Holotype; male; 64124329-7811-4df4-829c-f138766ad5c3; deposited at: Oxford University Museum of Natural History (OUM); Unknown
Phyllium bioculatum Gray, 1832 in Gray, 1835: 30.
... Show all ... (100)

Phyllium bioculatum Gray, 1832 in Audinet-Serville, 1838: 292.
Phyllium bioculatum Gray, 1832 in Burmeister, 1838: 590.
Phyllium bioculatum Gray, 1832 in Gray, 1843: 117.
Phyllium bioculatum Gray, 1832 in Westwood, 1859: 175.
Phyllium bioculatum Gray, 1832 in Stål, 1875: 105.
Phyllium bioculatum Gray, 1832 in Bolívar, 1895: 374.
Phyllium bioculatum Gray, 1832 in Rehn, 1904: 106.
Pulchriphyllium bioculatum (Gray, 1832) in Kirby, 1904: 421.
Phyllium bioculatum Gray, 1832 in Redtenbacher, 1906: 174.
Pulchriphyllium bioculatum (Gray, 1832) in Giglio-Tos, 1910: 14.
Phyllium bioculatum Gray, 1832 in Bolívar & Ferrière, 1912: 294. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Merle, 1917: 93.
Phyllium bioculatum Gray, 1832 in Chopard, 1920: 261, 263. Phylogeny or classification; Image or description
Phyllium bioculatum Gray, 1832 in Grimpe, 1921: 227. Life history; Behavior
Pulchriphyllium bioculatum (Gray, 1832) in Showalter, 1929: 28, fig..
Phyllium bioculatum Gray, 1832 in Rehn & Rehn, 1934: 411.
Pulchriphyllium bioculatum (Gray, 1832) in Orian, 1957: 517. Specimen or distribution
Pulchriphyllium bioculatum (Gray, 1832) in Vinson, 1968: 113.
Phyllium bioculatum Gray, 1832 in Klante, 1969: 2.
Phyllium bioculatum Gray, 1832 in Sellick, 1974: 27.
Phyllium bioculatum Gray, 1832 in Bedford, 1978: 126. Life history
Phyllium bioculatum Gray, 1832 in Robinson, 1980: 2.
Phyllium bioculatum Gray, 1832 in Sellick, 1980: 197. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Vanschuytbroeck & Cools, 1981: 25. Specimen or distribution
Pulchriphyllium bioculatum (Gray, 1832) in Passerin d' Entrèves, 1981: 55. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Hausleithner, 1983: 278.
Phyllium bioculatum Gray, 1832 in Brock, 1985: 28, fig. Image or description
Phyllium bioculatum Gray, 1832 in Carlberg, 1986: 6.
Phyllium bioculatum Gray, 1832 in Carlberg, 1986: 643. Life history
Phyllium bioculatum Gray, 1832 in Carlberg, 1987: 54. Life history
Phyllium bioculatum Gray, 1832 in Carlberg, 1987: 47. Structure
Phyllium bioculatum Gray, 1832 in Matyot, 1988: 11.
Phyllium bioculatum Gray, 1832 in Sellick, 1988: 280. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Ziegler, 1989: 30.
Phyllium bioculatum Gray, 1832 in Flückiger & Peltier, 1989: figs.
Phyllium bioculatum Gray, 1832 in Matyot, 1990: 32.
Phyllium bioculatum Gray, 1832 in Matyot, 1990: 4. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Preston-Mafham, 1990: 82. Behavior
Phyllium bioculatum Gray, 1832 in Matyot, 1992: 18.
Phyllium bioculatum Gray, 1832 in Brock, 1992: 13, 46, fig.
Phyllium bioculatum Gray, 1832 in Unno, 1993: 12-13, fig. Image or description
Phyllium bioculatum Gray, 1832 in Seow-Choen, Brock & Seow-En, 1994: 8.
Phyllium bioculatum Gray, 1832 in Schulten, 1995: 108, figs., pl. 7.
Phyllium bioculatum Gray, 1832 in Deschandol, 1995: 20.
Phyllium bioculatum Gray, 1832 in Ziegler, 1995: 32.
Phyllium bioculatum Gray, 1832 in Woolman & Dharmasiri, 1995: 33. Ecological data
Phyllium bioculatum Gray, 1832 in Hanibeltz, Nakamura, Imms & Abdullah, 1995: 60. Life history
Phyllium bioculatum Gray, 1832 in Schulten, 1995: 643, fig. 4.
Phyllium bioculatum Gray, 1832 in Brock, 1995: 98.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 1995: 25, fig. Image or description
Phyllium bioculatum Gray, 1832 in Tay & Seow-Choen, 1996: 183.
Phyllium bioculatum Gray, 1832 in Seow-Choen, Seow-En & Seow-An, 1996: 43.
Phyllium bioculatum Gray, 1832 in Seiler, 1996: 6.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1996: 72, figs..
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1997: 139, figs..
Phyllium bioculatum Gray, 1832 in Alderton, 1997: 17, fig. Image or description
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1998: 90.
Phyllium bioculatum Gray, 1832 in Brock, 1999: 23, 41, pl. 18b.
Phyllium bioculatum Gray, 1832 in Brock, 1999: 160, pl. 35-38, 185.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1999: 299.
Phyllium bioculatum Gray, 1832 in Seow-Choen, 1999: 74.
Phyllium bioculatum Gray, 1832 in Brock, 2000: 56, figs.
Phyllium bioculatum Gray, 1832 in Seiler, Bradler & Koch, 2000: 110, figs.
Phyllium bioculatum Gray, 1832 in Delfosse, 2000: 3.
Phyllium bioculatum Gray, 1832 in Zompro, 2000: 61. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Brock, 2000: 9. Image or description
Phyllium bioculatum Gray, 1832 in Bragg, 2001: 191, fig..
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Grösser, 2001: 53, figs.
Phyllium bioculatum Gray, 1832 in Baxter, 2002: 14, 91, figs.
Phyllium bioculatum Gray, 1832 in Alderton, 2002: 16, fig. Image or description
Phyllium bioculatum Gray, 1832 in Whiting, Bradler & Maxwell, 2003: 265.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 2003: 226, fig. 1, 231.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Brock, 2003: 42, fig. 24.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Zompro, 2004: 273. Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2005: 108.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Otte & Brock, 2005: 275.
Phyllium bioculatum Gray, 1832 in Sorpongpaisal & Thanasinchayakul, 2006: 22. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Bragg, 2007: 9. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Gerlach & Haas, 2008: 68. Image or description; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Grösser, 2008: 93, figs.. Image or description; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Hennemann, Conle, Gottardo & Bresseel, 2009: 80. Phylogeny or classification
Phyllium bioculatum Gray, 1832 in Narita, Pereira, Kjellberg & Kageyama, 2010: 74. Structure
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Mandal & Yadav, 2010: 53. Image or description
Phyllium bioculatum Gray, 1832 in Harman, 2011: 7. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Hutacharern, 2011: 639. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Harman, 2012: 12-13. Specimen or distribution
Phyllium bioculatum Gray, 1832 in Grösser, 2013: 46-51, figs.. Image or description; Phylogeny or classification; Specimen or distribution; Life history
Phyllium bioculatum Gray, 1832 in Bradler, 2015: 5, fig. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2016: 428, figs. Image or description
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2017: 153, figs. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2017: 202. Phylogeny or classification; Specimen or distribution
Phyllium bioculatum Gray, 1832 in Bradler & Buckley, 2018: 283, fig. Image or description
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Cumming, Le Tirant & Teemsma, 2018: 16. Image or description; Phylogeny or classification; Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Cumming & Le Tirant, 2018: 2.
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Laskar & Sinha, 2018: 1107-1111. Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2018: 666, figs. Image or description; Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, Seow-En, Chua & Choo, 2019: 382. Specimen or distribution
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Seow-Choen, 2020: 397. Specimen or distribution
Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Bank, Cumming, Li, Henze, Le Tirant & Bradler, 2021: Supp. Table 1. Phylogeny or classification
Phyllium (Pulchriphyllium) bioculatum Gray, 1832 in Singh, Chandra & Gupta, 2021: 228. Specimen or distribution

Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Seow-Choen, 2021: 899, figs. Phylogeny or classification
Pulchriphyllium bioculatum bioculatum (Gray, 1832) in Brock & Büscher, 2022: pl. 3-4 & 12, 425, 563, fig. Image or description; Specimen or distribution

Nomenclature references (104)

Alderton, D. (1997) Your first stick insect. Kingdom Books, Havant, 31 pp.
Alderton, D. (2002) How to Care for your Stick Insect. Kingdom Books, Havant, 34 pp.
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Bank, S., Cumming, R.T., Li, Y., Henze, K., Le Tirant, S. & Bradler, S. (2021) A tree of leaves: Phylogeny and historical biogeography of the leaf insects (Phasmatodea: Phylliidae). Communications Biology, 4(932), 1–12. Available at http://www.nature.com/articles/s42003-021-02436-z

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Bolívar, I. (1895) Mission scientifique de M. Ch. Alluaud aux íles Séchelles (Mars, Avril, May, 1892). Orthoptères. Appendix Orthoptères de la Réunion 6 mémoire. Annales de la Société Entomologique de France, 64, 372–375. Available at http://www.biodiversitylibrary.org/item/34283#page/382/mode/1up

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Brock, P.D. (1985) In The Phasmid Rearer’s Handbook (Stick Insects and Leaf Insects). The Amateur Entomologists’ Society. The Amateur Entomologist Vol. 20, Feltham. 41 pp.

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Descendants and synonyms

Stats

	Names
Rank	Total	Valid	Invalid
species	4	1	3