species

Dares ulula (Westwood, 1859)

Spiny Ulula Stick Insect

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female Acanthoderus ulula juv. (holotype). (CollectionObject).

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female Acanthoderus ulula juv., lateral view (holotype). (CollectionObject).

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female, head & thorax, lateral view of Acanthoderus ulula juv. (holotype). (CollectionObject).

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female juv., end of abdomen, dorsal view of Acanthoderus ulula (holotype). (CollectionObject).

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female juv., end of abdomen, lateral view of Acanthoderus ulula (holotype). (CollectionObject).

CollectionObject 1558300; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234: copyright UMO. female juv., end of abdomen, ventral view of Acanthoderus ulula (holotype). (CollectionObject).

CollectionObject 1558302; f0517d30-773d-4443-b1e6-8c08cdfa9611: copyright MNHN, Paris. male juv. of synonym Dares (Dares) corticinus (holotype). (CollectionObject).

CollectionObject 1558302; f0517d30-773d-4443-b1e6-8c08cdfa9611: copyright MNHN, Paris. male juv., lateral view of synonym Dares (Dares) corticinus (holotype). (CollectionObject).

Nomenclature (34)

Acanthoderus ulula Westwood, 1859: 53, pl. 26:1.
Holotype; immature; 3f27b43c-2ffb-45d4-b6ae-9ddb7caa2234; deposited at: Oxford University Museum of Natural History (OUM);
Dares ulula (Westwood, 1859) in Stål, 1875: 94.
... Show all ... (30)

Dares ulula (Westwood, 1859) in Kirby, 1904: 400.
Dares calamita Redtenbacher, 1906: 55.
Syntypes (n= 2); female; f18a25f5-7afd-4af0-81d4-fc1d67c26760; deposited at: Hungarian Natural History Museum , Budapest, Hungary (Termeszettudomanyi Muzeum) (HNHM); Unknown
Dares corticinus Redtenbacher, 1906: 56.
Holotype; immature; f0517d30-773d-4443-b1e6-8c08cdfa9611; deposited at: Museum National d'Histoire Naturelle, Paris (MNHN);
Dares (Dares) ulula (Westwood, 1859) in Redtenbacher, 1906: 54.
Dares ulula (Westwood, 1859) in Günther, 1932: 66.
Dares calamita Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Günther, 1935: 3.
Dares corticinus Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Günther, 1935: 3.
Dares ulula (Westwood, 1859) in Günther, 1935: 3.
Dares ulula (Westwood, 1859) in Abercrombie, 1992: 2.
Dares ulula (Westwood, 1859) in Bragg, 1993: 39.
Dares ulula (Westwood, 1859) in Abercrombie, 1995: 34.
Dares ulula (Westwood, 1859) in Bragg, 1995: 51. Life history
Dares ulula (Westwood, 1859) in Seow-Choen, 1997: 13, fig..
Dares calamita Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Bragg, 1998: 60.
Dares corticinus Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Bragg, 1998: 60.
Dares ulula (Westwood, 1859) in Sellick, 1998: 208. Phylogeny or classification
Dares ulula (Westwood, 1859) in Seow-Choen, 1998: 89.
Dares ulula (Westwood, 1859) in Bragg, 2001: 147, figs. 43-45.
Dares ulula (Westwood, 1859) in Brock, 2003: 69.
Dares corticinus Redtenbacher, 1906 in Zompro, 2004: 219.
Dares ulula (Westwood, 1859) in Zompro, 2004: 220. Phylogeny or classification
Dares ulula (Westwood, 1859) in Seow-Choen, Grinang & Naming, 2004: 394.
Dares calamita Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Otte & Brock, 2005: 118.
Dares corticinus Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Otte & Brock, 2005: 118.
Dares ulula (Westwood, 1859) in Otte & Brock, 2005: 118.
Dares ulula (Westwood, 1859) in Harman, 2012: 13. Specimen or distribution
Dares ulula (Westwood, 1859) in Seow-Choen, 2016: 380, figs. Image or description
Dares corticinus Redtenbacher, 1906 (subjective synonym of Acanthoderus ulula Westwood, 1859) in Delfosse, Cliquennois, Depraetere & Robillard, 2019: 196. Specimen or distribution
Dares ulula (Westwood, 1859) in Seow-Choen, 2019
Dares ulula (Westwood, 1859) in Brock & Büscher, 2022: 519. Specimen or distribution

Acanthoderus ulula Westwood, 1859 in Seow-Choen, 2025: 321. Specimen or distribution; Image or description
Dares ulula (Westwood, 1859) in Bragg, 2025: 85. Specimen or distribution

Nomenclature references (26)

Abercrombie, I. (1992) Observations on egg laying by Epidares nolimetangere (de Haan) and Dares ulula (Westwood). Phasmid Studies, 1(1), 2–4. Available at http://www.biodiversitylibrary.org/item/207247#page/4/mode/1up
Abercrombie, I. (1995) Beobachtungen bei der Eiablage bei Epidares nolimetangere (de Haan) und Dares ulula (Westwood). Arthropoda, 3(3), 34–36.
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Bragg, P.E. (1993) Parasites of Phasmida. The Entomologist, 112(1), 37–42. Available at http://www.researchgate.net/publication/349039160_Parasites_of_Phasmida

Bragg, P.E. (1995) The culture of Bornean phasmids. Phasmid Studies, 4(2), 50–53. Available at http://www.biodiversitylibrary.org/item/207262#page/52/mode/1up

Bragg, P.E. (1998) A revision of the Heteropteryginae (Insecta: Phasmida: Bacillidae) of Borneo, with the description of a new genus and ten species. Zoologische Verhandlingen, 316, 1–135. Available at http://www.repository.naturalis.nl/document/148998

Bragg, P.E. (2001) Phasmids of Borneo. Natural History Publications, Kota Kinabalu, Sabah, 771 pp. Available at http://www.nhpborneo.com/book/phasmids-of-borneo/

Bragg, P.E. (2025) A review of the cockroaches, mantids and phasmids from Niah national park, Sarawak (Insecta: Blattodea, Mantodea, and Phasmida). The Sarawak Museum Journal, 89(110), 73–91.

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. (2003) Rearing and studying stick and leaf insects. Revised Edition. The Amateur Entomologists' Society, London, Vol. 22, 89 pp. Available at http://shop.amentsoc.org/rearing-and-studying-stick-and-leaf-insects.html

Delfosse, E., Cliquennois, N., Depraetere, M. & Robillard, T. (2019) Catalogue des types de la collection de phasmes du Muséum national d’Histoire naturelle de Paris (Insecta, Phasmatodea). Zoosystema, 41(11), 181–235. Available at http://sciencepress.mnhn.fr/en/periodiques/zoosystema/41/11

Günther, K. (1932) Die von Professor Dr. Winkler 1924/25 in Zentralborneo gesammelten Phasmoiden. Zoologischer Anzeiger, 101(3), 65–73.

Günther, K. (1935) Phasmoiden aus Centralborneo. Gesammelt von Dr. Mjöberg 1925-26. Arkiv för Zoologi, (9), 1–29.

Harman, A.J.E. (2012) The Development of the Phasmid Species List Part Three. PSG No. 101 - PSG No. 150. The Phasmid Study Group Newsletter, 128, 13–15.

Kirby, W.F. (1904) A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). The Trustees of the British Museum, London, Vol. 1, 501 pp. Available at http://www.biodiversitylibrary.org/item/29661

Otte, D. & Brock, P.D. (2005) In Phasmida Species File. Catalog of Stick and Leaf Insects of the world. The Insect Diversity Association at the Academy of Natural Sciences, Philadelphia. 414 pp. Available at http://www.goodreads.com/book/show/24725959-phasmida-species-file

Redtenbacher, J. (1906) Phasmidae Areolatae. In Die Insektenfamilie der Phasmiden. Vol. 1, 1–180, pls 1–6. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmidenVol.1

Sellick, J.T.C. (1998) The micropylar plate of the eggs of Phasmida, with a survey of the range of plate form within the order. Systematic Entomology, 23, 203–228. Available at http://onlinelibrary.wiley.com/doi/abs/10.1046/j.1365-3113.1998.00056.x

Seow-Choen, F. (1997) The joy of stick-insects. Singapore Scientist, 81, 10–14. Available at http://www.researchgate.net/profile/Francis-Seow-Choen

Seow-Choen, F. (1998) List of stick-insects (Phasmida) in the collection of the Centre for Insect Systematics Universiti Kebangsaan Malaysia. Serangga, 3(1), 87–92. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Seow-Choen, F. (2016) A taxonomic guide to the stick insects of Borneo. Natural History Publications (Borneo), Kota Kinabalu, 454 pp. Available at http://www.nhpborneo.com/book/taxonomic-guide-stick-insects-borneo/

Seow-Choen, F. (2019) A taxonomic guide to the stick insects of Borneo: Vol. III. Natural History Publications (Borneo), Kota Kinabalu, 278 pp. Available at http://www.nhpborneo.com/book/a-taxonomic-guide-to-the-stick-insects-of-borneo-vol-3/

Seow-Choen, F. (2025) A Photographic Guide: Stick Insects of Borneo. Natural History Publications (Borneo), 387 pp.

Seow-Choen, F., Grinang, J. & Naming, M. (2004) Phasmids. In Sarawak Bau Limestone Biodiversity. Sarawak Museum Department. Sarawak Museum Journal, Volume 59, No 80, Kuching. pp. 391–401. Available at http://www.researchgate.net/profile/Francis_Seow-Choen

Stål, C. (1875) Recensio orthopterorum. Revue critique des Orthoptères décrits par Linné, DeGeer et Thunberg. 3. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 32, 1–105. Available at http://hdl.handle.net/2027/coo.31924018284798

Westwood, J.O. (1859) Catalogue of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. British Museum, London, 196 pp, 48 pl. Available at http://www.archive.org/details/catalogueofortho00brituoft
Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Goecke & Evers, Keltern-Weiler, Germany, 327 pp. Available at http://www.researchgate.net/publication/313233330_Revision_of_the_genera_of_the_Areolatae_including_the_status_of_Timema_and_Agathemera_Insecta_Phasmatodea

Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
species	1	0	4	5