genus

Clonaria Stål, 1875

Type

Bacillus natalis Westwood, 1859 type species by subsequent designation of Clonaria Stål, 1875

Nomenclature (55)

Clonaria Stål, 1875: 12, 14, 71. (valid)
type species by subsequent designation Bacillus natalis Westwood, 1859 in Kirby, 1904
Clonaria Stål, 1875: 5, 13.
... Show all ... (51)

Clonaria Stål, 1875: 9. Phylogeny or classification
Gratidia Stål, 1875: 14, 70.
type species by original monotypy Gratidia sansibara Stål, 1875
Clonaria Stål, 1875 in Brunner von Wattenwyl, 1893: 89.
Paraclonaria Brunner von Wattenwyl, 1893: 89.
type species by subsequent designation Paraclonaria hamuligera Schulthess, 1898 in Rehn, 1904
Gratidia Stål, 1875 in Brunner von Wattenwyl, 1893: 88. Phylogeny or classification
Gratidia Stål, 1875 in Karsch, 1898: 365. Phylogeny or classification
Clonaria Stål, 1875 in Kirby, 1904: 337.
Paraclonaria Brunner von Wattenwyl, 1893 in Rehn, 1904: 83.
Paraclonaria Brunner von Wattenwyl, 1893 in Kirby, 1904: 335.
Gratidia Stål, 1875 in Kirby, 1904: 330. Phylogeny or classification
Gratidia Stål, 1875 (subjective synonym of Clonaria Stål, 1875) in Rehn, 1904: 82.
Gratidia Stål, 1875 in Brunner von Wattenwyl, 1907: 217. Phylogeny or classification
Paraclonaria Brunner von Wattenwyl, 1893 (subjective synonym of Gratidia Stål, 1875) in Brunner von Wattenwyl, 1907: 217. Phylogeny or classification
Gratidia Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Karny, 1923: 236. Phylogeny or classification
Clonaria Stål, 1875 in Rehn, 1933: 63.
Gratidia Stål, 1875 (subjective synonym of Clonaria Stål, 1875) in Rehn, 1933: 63.
Gratidia Stål, 1875 in Ebner, 1934: 219. Phylogeny or classification
Gratidia Stål, 1875 in Mishchenko, 1941: 147. Phylogeny or classification
Gratidia Stål, 1875 in Bey-Bienko, 1946: 75. Phylogeny or classification
Gratidia Stål, 1875 in Kevan, 1950: 200. Phylogeny or classification
Clonaria Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Günther, 1957: 92. Phylogeny or classification
Gratidia Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Günther, 1957: 87, 92. Phylogeny or classification
Gratidia Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Bradley & Galil, 1977: 185. Phylogeny or classification
Clonaria Stål, 1875 in James, 1982: 7.
Clonaria Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Schulten, 1995: 64. Image or description
Gratidia Stål, 1875 in Bragg, 1995: 12. Phylogeny or classification
Gratidia Stål, 1875 in Fritzsche, 1996: 13. Specimen or distribution
Gratidia Stål, 1875 in Sellick, 1997: 116. Phylogeny or classification
Gratidia Stål, 1875 in Sellick, 1998: 218. Phylogeny or classification
Gratidia Stål, 1875 in Brock, 1998: 26. Phylogeny or classification
Clonaria Stål, 1875 (subjective synonym of Ramulus Saussure, 1862) in Lipinski, Greven, Schulten & Löser, 1999: 54. Image or description
Gratidia Stål, 1875 in Bragg, 2001: 633. Phylogeny or classification
Paraclonaria Brunner von Wattenwyl, 1893 (subjective synonym of Gratidia Stål, 1875) in Bragg, 2001: 625. Phylogeny or classification
Clonaria Stål, 1875 in Zompro, 2004: 308.
Paraclonaria Brunner von Wattenwyl, 1893 in Zompro, 2004: 317.
Gratidia Stål, 1875 in Zompro, 2004: 311. Phylogeny or classification
Clonaria Stål, 1875 in Cliquennois, 2005: 12.
Clonaria Stål, 1875 in Hanot, 2005: 15.
Clonaria Stål, 1875 in Brock, 2005: 26.
Clonaria Stål, 1875 in Otte & Brock, 2005: 97.
Gratidia Stål, 1875 (subjective synonym of Clonaria Stål, 1875) in Brock, 2005: 26.
Gratidia Stål, 1875 (subjective synonym of Clonaria Stål, 1875) in Otte & Brock, 2005: 97.
Paraclonaria Brunner von Wattenwyl, 1893 (subjective synonym of Clonaria Stål, 1875) in Otte & Brock, 2005: 97.
Clonaria Stål, 1875 in Brock, 2006: 41.
Clonaria Stål, 1875 in Mandal & Yadav, 2010: 21. Image or description
Clonaria Stål, 1875 in Harman, 2011: 11. Specimen or distribution
Clonaria Stål, 1875 in Harman, 2012: 13, 14. Specimen or distribution
Clonaria Stål, 1875 in Feulner, 2012: 77. Image or description; Specimen or distribution
Clonaria Stål, 1875 in Harman, 2013: 13. Specimen or distribution
Clonaria Stål, 1875 in Bidau, Taffarel & Castillo, 2016: 29. Behavior
Clonaria Stål, 1875 in Bradburne, 2019: 30, figs. Image or description; Ecological data

Clonaria Stål, 1875 in Brock & Büscher, 2022: 506. Specimen or distribution
Clonaria Stål, 1875 in Mirzaee, Büscher, Evangelista, Eggleton, Anisyutkin & Legendre, 2026: 383. Specimen or distribution

Nomenclature references (41)

Bey-Bienko, G.Y. (1946) A new species of the genus Gratidia Stål (Phasmoidea) from Iran, with a list of palaearctic species of this genus. Proceedings of the Royal Entomological Society, London, 15(5-6), 74–76. Available at http://onlinelibrary.wiley.com/doi/10.1111/j.1365-3113.1946.tb00826.x
Bidau, C.J., Taffarel, A. & Castillo, E.R. (2016) Breaking the rule: multiple patterns of scaling of sexual size dimorphism with body size in orthopteroid insects. Revista de la Sociedad Entomológica Argentina, 75(1-2), 11–36. [2015] Available at http://www.researchgate.net/publication/304497290_Breaking_the_rule_multiple_patterns_of_scaling_of_sexual_size_dimorphism_with_body_size_in_orthopteroid_insects
... Show all ... (37)

Bradburne, R.P. (2019) Three new localities for proposed Clonaria sp. phasmids in Southern Uganda. Phasmid Studies, 20, 30–38. Available at http://phasmidstudygroup.org/files/Phasmid_Studies/Phasmid_Studies_Vol20.pdf

Bradley, J.C. & Galil, B.S. (1977) The taxonomic arrangement of the Phasmatodea with keys to the subfamilies and tribes. Proceedings of the Entomological Society, Washington, 79(2), 176–208. Available at http://www.biodiversitylibrary.org/page/16301624#page/194/mode/1up

Bragg, P.E. (1995) Type species of phasmid genera with particular reference to the status of Baculum Saussure, 1861, Ramulus, Saussure, 1862, and Gratidia Stål, 1875. Phasmid Studies, 4(1), 11–14. Available at http://www.biodiversitylibrary.org/item/207262#page/13/mode/1up

Bragg, P.E. (2001) Phasmids of Borneo. Natural History Publications, Kota Kinabalu, Sabah, 771 pp. Available at http://www.nhpborneo.com/book/phasmids-of-borneo/

Brock, P.D. & Büscher, T.H. (2022) Stick and leaf-insects of the world, Phasmids. NAP Editions, France: Verrières-le-Buisson, 612 pp. Available at http://www.napeditions.com/en/67-stick-and-leaf-insects-of-the-world.html

Brock, P.D. (1998) Type material of stick-insects (Insecta: Phasmida) in the Zoological Museum, University of Copenhagen. Steenstrupia, 24(1), 23–35.

Brock, P.D. (2005) A new genus and four new species of South African stick insects. Phasmid Studies, 12(1-2), 25–37. Available at http://www.biodiversitylibrary.org/item/207169#page/27/mode/1up

Brock, P.D. (2006) Three new species of South African stick insects (Phasmida). Journal of Orthoptera Research, 15(1), 37–44. Available at http://bioone.org/journals/journal-of-orthoptera-research/volume-15/issue-1/1082-6467(2006)15[37:TNSOSA]2.0.CO;2/Three-new-species-of-South-African-stick-insects-Phasmida/10.1665/1082-6467(2006)15[37:TNSOSA]2.0.CO;2.full

Brunner von Wattenwyl, C. (1893) Révision du Système des Orthoptères et description des espèces rapportées par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova, (2)13(33), 1–230. Available at http://www.biodiversitylibrary.org/bibliography/5121

Brunner von Wattenwyl, C. (1907) Phasmidae Anareolatae (Clitumnini, Lonchodini, Bacunculini). In Die Insektenfamilie der Phasmiden: Vol. II. Verlag Engelmann, Leipzig. 181–338, pls 7–15. [in German] Available at http://www.archive.org/details/DieInsektenfamilieDerPhasmiden.Vol.Ii.PhasmidaeAnareolataeclitumnini

Cliquennois, N. (2005) À propos des Gratidiini: notes critiques (Phasmatodea, Anareolatae. Bulletin de Phyllie, 22, 12–28. [2004] Available at http://https://www.researchgate.net/profile/Nicolas_Cliquennois

Ebner, R. (1934) Wissenschaftliche Ergebnisse der mit der Unterstützung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-ägyptischen Sudan (Kordofan) 1914. 28. Orthoptera B. (Dermaptera, Blattidae, Phasmidae). Denkschriften der Akademie der Wissenschaften in Wien Mathematisch-Naturwissenschaftlische Klasse, 103, 211–222. [1933] Available at http://www.zobodat.at/pdf/DAKW_103_0211-0222.pdf

Feulner, G.R. (2012) Stick Insects (Order Phasmida) of Eastern Arabia: A Photographic Review. Tribulus: Bulletin of the Emirates Natural History Group, 20, 77–82. Available at http://www.enhg.org/Portals/1/trib/V20/TribulusV20.pdf

Fritzsche, I. (1996) A report on Gratidia sp. from Zaire (PSG 141), and a study of the hatching of the eggs. Phasmid Studies, 5(1), 13–17. Available at http://www.biodiversitylibrary.org/item/207389#page/15/mode/1up

Günther, K. (1957) Chapter III Phasmatoptera. In B. Hanstrom, P. Brinck, & G. Rudebeck (Eds.). South African Animal life, results of the Lund University expedition in 1950-51. Almqvist & Wiksell, Stockholm.

Hanot, C. (2005) Comportement de ponte chez Clonaria Stal, 1875 sp. (Insecta: Phasmatodea, Pachymorphinae) [Egg laying behaviour in Clonaria Stal, 1875 sp. (Insecta: Phasmatodea, Pachymorphinae)]. Le Bulletin d'Arthropoda, 25, 15–16.

Harman, A.J.E. (2011) The Development of the Phasmid Species List Part One: PSG No. 1-50. The Phasmid Study Group Newsletter, 125, 6–13. Available at http://phasmidstudygroup.org/files/PSG_Newsletters/Newsletter_125.pdf

Harman, A.J.E. (2012) The Development of the Phasmid Species List Part Three. PSG No. 101 - PSG No. 150. The Phasmid Study Group Newsletter, 128, 13–15.

Harman, A.J.E. (2013) The Development of the Phasmid Species List Part Five: PSG No.201 - PSG No. 250. The Phasmid Study Group Newsletter, 130, 13–15. Available at http://phasmidstudygroup.org/files/PSG_Newsletters/Newsletter_130.pdf

James, T. (1982) PSG/42 TA U3. The Phasmid Study Group Newsletter, 9/10, 7–8.

Karny, H.H. (1923) Zur Nomenklatur der Phasmoiden. Hortus Botanicus Bogoriensis, Java (Treubia), 3(2), 230–242. Available at http://www.biodiversitylibrary.org/item/110723#page/310/mode/1up

Karsch, F.A.F. (1898) Vorarbeiten zu einer Orthopterologie Ostafrika's. I. Die Gespenstheuschrecken, Phasmodea. Entomologische Nachrichten, 24, 359–383. Available at http://www.biodiversitylibrary.org/item/42365#page/373/mode/1up

Kevan, D.K.M.E. (1950) Orthoptera from the hills of south-east Kenya. Journal of the East Africa Natural History Society, 19(5), 192–221. Available at http://www.biodiversitylibrary.org/content/part/EANHS/XIX_No.5_89__192_1947-1948_Kevan.pdf

Kirby, W.F. (1904) A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). The Trustees of the British Museum, London, Vol. 1, 501 pp. Available at http://www.biodiversitylibrary.org/item/29661

Lipinski, K., Greven, H., Schulten, D. & Löser, S. (1999) In Die Struktur der Eihüllen von 48 Phasmatodea-Arten aus der Sammlung des Löbbecke Museum und Aquazoo Düsseldorf. Entomologische Mitteilungen aus dem Löbbecke-Museum und Aquazoo. 1–125 pp. [in German] Available at http://www.zobodat.at/pdf/Entomologie-heute_BH_5_0001-0125.pdf

Mandal, S.K. & Yadav, K. (2010) Some phasmida (stick and leaf insects) of India. RECORDS OF THE ZOOLOGICAL SURVEY OF INDIA OCCASIONAL PAPER, 318, 1–64. Available at http://faunaofindia.nic.in/PDFVolumes/occpapers/318/index.pdf

Mirzaee, Z., Büscher, T.H., Evangelista, D.A., Eggleton, P., Anisyutkin, L.N. & Legendre, F. (2026) Arthropod diversity in Iran: Class Insecta–Polyneoptera, part I. Blattodea Brunner von Wattenwyl, 1882, Mantodea Burmeister, 1838, and Phasmatodea Jacobson & Bianchi, 1902. Journal of Insect Biodiversity and Systematics, 12(2), 357–394. Available at https://www.researchgate.net/publication/401317940_Arthropod_diversity_in_Iran_Class_Insecta-Polyneoptera_part_I_Blattodea_Brunner_von_Wattenwyl_1882_Mantodea_Burmeister_1838_and_Phasmatodea_Jacobson_Bianchi_1902

Mishchenko, L. (1941) The species of the genus Gratidia Stål in USSR (Phasmidae, Clitumnidae). Mitteilungen der Akademie der Wissenschaften der Georgischen SSR, 2 (1-2), 145–148.

Otte, D. & Brock, P.D. (2005) In Phasmida Species File. Catalog of Stick and Leaf Insects of the world. The Insect Diversity Association at the Academy of Natural Sciences, Philadelphia. 414 pp. Available at http://www.goodreads.com/book/show/24725959-phasmida-species-file

Rehn, J.A.G. (1904) Studies in the Orthopterous family Phasmidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 56, 38–107. Available at http://www.biodiversitylibrary.org/item/30096#page/48/mode/1up

Rehn, J.A.G. (1933) Dermaptera and Orthoptera of the De Schauensee South African Expedition, - Part I. Proceedings of the Academy of Natural Sciences of Philadelphia, 85, 39–51, pl. 1. Available at http://www.jstor.org/stable/4064168?refreqid=excelsior%3A5a00cb35a8814d17c5975d8eef3d4d18&seq=1

Saussure, H.L.F.de (1862) Etudes sur quelques orthoptères du musée de Genève nouveaux ou imparfaitement connus. Annales de la Société Entomologique de France, (4)1, 469–494, pls. 11–12. [1861] Available at http://www.biodiversitylibrary.org/item/34164#page/477/mode/1up

Schulten, D. (1995) In Wandelnde Blätter, Stab- und Gespenstschrecken. Entomologische Mitteilungen aus dem Löbbecke-Museum and Aquazoo. Vol. 3, 1–132 pp. [in German] Available at http://www.zobodat.at/pdf/Entomologie-heute_BH_3_0001-0132.pdf

Sellick, J.T.C. (1997) Descriptive terminology of the phasmid egg capsule, with an extended key to the phasmid genera based on egg structure. Systematic Entomology, 22, 97–122. Available at http://onlinelibrary.wiley.com/doi/abs/10.1046/j.1365-3113.1997.d01-30.x

Sellick, J.T.C. (1998) The micropylar plate of the eggs of Phasmida, with a survey of the range of plate form within the order. Systematic Entomology, 23, 203–228. Available at http://onlinelibrary.wiley.com/doi/abs/10.1046/j.1365-3113.1998.00056.x

Stål, C. (1875) Observations Orthoptérologiques. 1. Sur une systématisation nouvelle des Phasmides. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 3(14), 1–21. Available at https://www.biodiversitylibrary.org/item/49924#page/523/mode/1up

Stål, C. (1875) Recensio orthopterorum. Revue critique des Orthoptères décrits par Linné, DeGeer et Thunberg. 3. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 32, 1–105. Available at http://hdl.handle.net/2027/coo.31924018284798

Stål, C. (1875) Recherches sur le Système des Phasmides. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 2(17), 1–19. Available at http://www.biodiversitylibrary.org/item/49927#page/573/mode/1up
Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea). Goecke & Evers, Keltern-Weiler, Germany, 327 pp. Available at http://www.researchgate.net/publication/313233330_Revision_of_the_genera_of_the_Areolatae_including_the_status_of_Timema_and_Agathemera_Insecta_Phasmatodea

Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon	Valid names
Rank	Extant	Fossil	Invalid	Total
genus	1	0	2	3
species	127	0	200	327